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Eurasian beaver (Castor fiber) sits up, captive, Devon, UK, May
Arctic Fox (Vulpes lagopus), in winter coat portrait, Svalbard, Norway, April
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Dolgoch Falls, Gwynedd, Wales, UK, May 2015
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
Nurse sharks (Ginglymostoma cirratum) over seagrass at night. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Pallass cat (Otocolobus manul) two sitting side by side, Tibetan Plateau, Qinghai, China
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) Mindo, Pichincha, Ecuador
Black and white view of Plains Zebra (Equus quagga) herd, Etosha National Park, Kunene, Namibia
Aerial shot of the Laguna Sian Ka an, freshwater lagoon, at edge of the sea, close to Boca Paila, Quintana Roo, Yucatan Peninsula, Mexico. Photographed for The Freshwater Project February 2014
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
Buachaille Etive Beag reflected in Lochan na Fola after snowfall, early morning light, Glencoe, Scotland, UK. March 2017
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Norfolk Island pine trees (Araucaria heterophylla) near lagoon with Mount Lidgbird and Mount Gower in the background, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Commensal emperor shrimp (Periclimenes imperator) hitching a ride on a Nudibranch (Risbecia tryoni). Lembeh Strait, North Sulawesi, Indonesia
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Waterwheel plant (Aldrovanda vesiculosa) a carnivorous aquatic plant with Mosquito larva
Millstones made from millstone grit beneath Stannage Edge, Hathersage, Derbyshire, November 2016
Deserta Grande wolf spider (Hogna ingens), Deserta Grande, Madeira, Portugal. Critically endangered
Deep sea jellyfish (Periphylla periphylla), Trondheimsfjord, Norway, July
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Fin whale (Balaenoptera physalus) just below surface, south Barcelona coast, Spain, Mediterranean Sea
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Blue British Shorthair cat lying on his back
Silver tabby cat, Loki, age 7 months, walking with tail up
Border Terrier puppy, age 5 weeks, with raised paw
Ramparts of the prehistoric hill fort on Hambledon Hill above the Blackmore Vale, Dorset, England, UK, September 2015
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Flowers (Trollius sp) Balang Mountain, Wolong National Nature Reserve, Sichuan Giant Panda Sanctuaries - Wolong, Mt Siguniang and Jiajin Mountains UNESCO World Heritage Site, Sichuan Province, China
Hammerhead worms (Bipalium sp) on leaf, Sarawak. Borneo
Whinchat (Saxicola rubetra), male perched, Finland, May
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Saffron crocuses (Crocus sativus), cultivated for saffron, Lleida, Catalonia, Spain, November
The Senate Group of Giant sequoia (Sequoiadendron giganteum) trees on the Congress Trail in Sequoia National Park, California, USA
Common European toad tadpoles (Bufo bufo), sequence showing development of limbs. Derbyshire, UK. August. Digital composite
Soft corals (Dendronephthya sp) in coral reef, West Papua, New Guinea
Adult Eyelash Pit Viper or Eyelash Viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form
Meerkats (Suricata suricatta) with young, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
White rhino (Ceratotherium simum) at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Close up view of Honey Bee comb showing larvae in cells Norfolk, England, June 2017
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Peony (Paeonia officinalis) Mt Vettore, Umbria, Italy, June
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island
Seychelles Sunbird (Cinnyris dussumieri), adult male singing, Praslin Island, Republic of Seychelles
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Palm spider (Nephila inaurata) in its web, female, Praslin Island, Republic of Seychelles
Grey wolf (Canis lupus), adult with cub, captive, USA
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Fin whale (Balaenoptera physalus) with ctenophore in front of its mouth, south Barcelona coast, Spain, Mediterranean Sea
Jaguar (Panthera onca) female snarling, native to Southern and Central America, captive
Norfolk Terrier dog, Otto, age 1 year
English Oak (Quercus robur) in arable field with wheat, composite image showing all four seasons, Surrey, England, UK
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Conifer trees in mist at Alpe de Lerosa, Dolomite Mountains, Belluno Province, Veneto, Italy, September 2015
Thomass Langur (Presbytis thomasi), animal endemic to northern Sumatra. These primates have relatively long arms and legs, adapted for an acrobatic life in the crowns of trees
Lowland Streaked Tenrec (Hemicentetes semispinosus) active on forest floor at night. Masoala National Park, north east Madagascar
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February