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American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
Dhole / Indian wild dog {Cuon alpinus} Kanha NP, India
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Madagascar pitcher plants {Nepenthes madagascariensis} Pangalanes canal, East Madagascar
Madagascan reed frog {Heterixalus madagascar- iensis} Masoala NP, Madagascar
Painted mantella frog (Mantella madagascariensis) Mantadia NP, Eastern Madagascar
Bornean horned frog (Megophrys nasuta) on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Fosa of Fossa (Cryptoprocta ferox) male walking through dry deciduous forest, Kirindy, western Madagascar
Red fox (Vulpes vulpes) adult hunting for rodents by snow diving in deep snow. Hayden Valley, Yellowstone, USA. January
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Puma (Puma concolor puma), female sitting in grassland. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. November
Eyelash pit viper (Bothriechis schlegelii) (Viperidae: Crotalinae). Distictive yellow / orange oropel form. Arboreal species resting on wild ginger flower in mid-altitude rainforest under storey
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Net-casting / Ogre-faced Spider (Deinopis sp. ) with net. Active at night, Masoala National ParkNet-casting / Ogre-faced Spider (Deinopis sp.) with net. Active at night, Masoala National Park, Madagascar
Malayan colugo / Flying lemur (Cynocephalus variegatus) feeding on algae on tree bark. Danum Valley, Sabah, Borneo, Malaysia
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Aye-aye (Daubentonia madagascariensis) adult active and foraging in forest canopy at night. Deciduous forests, Daraina, northern Madagascar. Endangered endemic species
Boky-boky (Mungotictis decemlineata) or Narrow-striped mongoose adult foraging in leaf litter. Dry deciduous forest, Kirindy, western Madagascar
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Floating leaves of Giant water lily (Victoria amazonica). Northern Pantanal, Cuiaba River, Mato Grosso, Brazil
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Lined Leaf-tailed Gecko (Uroplatus lineatus). Active in dead palm fronds in forest understorey at night. Marojejy National Park, Madagascar
RF - Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered species
North American Bobcat (Lynx rufus) stalking along the edge of the Madison River. Yellowstone National Park, Wyoming, USA. January
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Dumerils Boa (Acrantophis dumerili) basking in late afternoon sunshine on forest floor. Berenty Private Reserve, southern Madagascar
Verreauxs sifaka lemur (Propithecus verreauxi) dancing or skipping across open ground, Gallery Forest, Berenty Reserve, southern Madagascar
Large aerial pitchers of natural hybrid Pitcher Plant (Nepenthes stenophylla x Nepenthes veitchi). Montane mossy heath forest or kerangas, Maliau Basin, Sabahs Lost World, Borneo
Hyacinth Macaw (Anodorhynchus hyacinthinus) in forest bordering of the Cuiaba River, Northern Pantanal, Brazil. September
Grandidiers / Giant Striped Mongoose (Galidictis grandidieri) foraging at night. Western edge of the Mahafaly Plateau, Tsimanampetsotsa National Park, south west Madagascar
Male Panther Chameleon (Furcifer pardalis) stalking prey in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
Juvenile Reticulated python (Python reticulatus) resting coiled round sapling by a stream. Danum Valley, Sabah, Borneo, Malaysia
Close up of juvenile Reticulated python (Python reticulatus) looking down from tree, Danum Valley, Sabah, Borneo, Malaysia
Flat-headed cat (Prionailurus planiceps) hunting frogs at night. Menanggol River, Sabah, Borneo
Toco toucan (Ramphastos toco) acrobatically perched, feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil
Jaguar (Panthera onca) female running along a sand spit chasing after a caiman. Northern Pantanal Cuiaba River, Mato Grosso, Brazil
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Sulawesi bear cuscus or Sulawesi bear phalanger (Ailurops ursinus) adult in forest canopy, showing use of prehensile tail. Tangkoko National Park, Sulawesi, Indonesia
North American river otter (Lutra canadiensis) two on frozen river edge. Upper Yellowstone River, Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Swallowtail butterfly (Papilio machaon) nectaring on Meadow clary (Salvia pratensis). North Tyrol, Austria. June
Alpine Wildflower meadow, a diversity of species including Meadow clary (Salvia pratensis), Sainfoin (Onobrychis sp) and Rattle (Rhinanthus sp). North Tyrol, Austria. June
Western red panda (Ailurus fulgens fulgens) climbing in tree. Singalila National Park, India / Nepal border
Alpine rose (Rhododendron ferrugineum) flowering beside mountain stream. North Tyrol, Austria. June
Coyote (Canis latrans) foraging in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
RF - Skull of a male Himalayan ibex (Capra sibirica) (killed by a snow leopard) lying on steep rocky slope. Himalayas, Ladakh, northern India
RF - Wild Snow leopard (Panthera uncia) stalking prey over broken rocky terrain. Ladakh Range, Western Himalayas, Ladakh, India
American Bison (Bison bison) frost covered male, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
Wild snow leopard (Panthera uncia) bounding down steep rock face in pursuit of prey. Ladakh Range, Western Himalayas, Ladakh, India
Silky sifaka (Propithecus candidus) female with baby sitting amongst branches in understorey of mid-altitude montane rainforest. Marojejy National Park, north east Madagascar
Red-and-green macaw (Ara chloropterus) flock feeding at wall of clay lick, flying and perched in trees. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Leopard (Panthera pardus) female climbing an Acacia tree, Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
Leopard (Panthera pardus) female with one of its cubs, killing a feral / domestic dog on rocky outcrop. Jawai / Bera, Rajasthan, India
Preening red-and-green macaw or green-winged macaw (Ara chloropterus) (Family Psittacidae). Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Red-and-green macaws (Ara chloropterus) pair in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Eyelash Viper (Bothriechis schlegelii) Distictive yellow / orange oropel form. Arboreal species resting in mid-altitude rainforest under storey. Caribbean slope, Costa Rica, Central America
Adult King Vulture (Sarcoramphus papa) feeding at a carcass. Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
RF - Daraina Sportive Lemur (Lepilemur milanoii) in day time rest tree hole in dry forest, northern Madagascar
RF - Male Violet Sabrewing hummingbirds (Campylopterus hemileucurus) hovering in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica
Growling riflebird (Ptiloris intercedens) male, Adelbert Mountain Range, Madang Province, Papua New Guinea. Endemic
Magnificent Bird of Paradise (Diphyllodes magnificus) male, Lower montane highlands near Mount Hagen, Enga Province, Papua New Guinea. Endemic
Waglers pit viper (Tropidolaemus wagleri) basking in mist-shrouded forest under storey. Danum Valley, Sabah, Borneo
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
Group of Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Bornean sun bear (Helarctos malayanus euryspilus) climbing tree at Bornean Sun Bear Conservation Centre (BSBCC), Sepilok, Sabah, Borneo
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at dawn. Kasanka National Park, Zambia
Female Diademed Sifaka (Propithecus diadema) with 2-month infant in the forest canopy. Andasibe-Mantadia National Park, eastern Madagascar. Endangered
