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Marsh-marigold (Caltha palustris) watercolour painting
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Mute swan (Cygnus olor) stretching its wings after preening on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Common starling (Sturnus vulgaris) murmuration, flock pursued by Peregrine falcon (Falco peregrinus) before landing at winter roost. The Netherlands. February 2020
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Spiny dogfish (Squalus acanthias) portrait. Rhode Island, New England, USA. August
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Southern elephant seal (Mirounga leonina) beach master / bull roaring amongst females and pups, in breeding colony. St Andrews Bay, South Georgia. October
Western hemlock (Tsuga heterophylla) in Douglas-fir forest (Pseudotsuga menziesii), Oregon, USA, June
Taiwan Blue Magpie, (Urocissa caerulea) endemic species, Taipei city park, Taiwan
Taiwan yuhina (Yuhina brunneiceps) perched amongst pink blossom, Alishan National Scenic Area, Taiwan. Endemic species
Large leaf of Giant Elephants Ear (Alocasia odora), in rainforest, Yangminshan, Taiwan
Japanese bullhead shark (Heterodontus japonicus) on reef. Chiba Prefecture, Honshu, Japan. May
Portrait of a Victorian koala (Phascolarctos cinereus). Koalas from the more southern and colder states of Australia have longer hair than there more northly counterparts
Arrowleaf balsamroot (Balsamorhiza sagittata) and lupins (Lupinus sp.), Columbia River Gorge, Oregon, USA. April
Lowland copperhead snake (Austrelaps superbus) male basking on rusty trailer, on farm in evening. Melbourne, Victoria, Australia
Dotterel, (Charadrius morinellus) female amongst Blue mountain heath (Phyllodoce caerulea) flowers, Ljungdalsfjallen, Harjedalen, Sweden
Male Zander or Pikeperch (Stizostedion / Sander lucioperca) with mouth wide open, defending nest. Lake di Lugano or Ceresio, Ticino, Switzerland
Alpine ibex (Capra ibex) resting in mountain pasture, Ticino, Switzerland
Loggerhead turtle (Caretta caretta) accompanied by pilotfish (Naucrates ductor) swimming near the surface. Vulnerable. Santa Maria Island, Azores, Portugal, Atlantic Ocean
Adriatic sturgeon (Acipenser naccarii) Parco del Ticino, Biosphere Reserve, Lombardia, Italy. Captive. Critically endangered species
Atlantic spotted dolphins, (Stenella frontalis), Formigas Islet dive site, Santa Maria Island, Azores, Portugal, Atlantic Ocean
Saddle-billed stork (Ephippiorhynchus senegalensis) flapping wings after preening. Liuwa Plain National Park, Zambia
African fish eagle (Haliaeetus vocifer) swoops to catch a freshly caught fish, dropped by a Saddle-billed stork (Ephippiorhynchus senegalensis) after being pressurised to do so by the eagle
Banyan fig tree (Ficus benjamina), one single individual tree, Banyan garden protected forest, Kenting National Park, Taiwan
Golden Birdwing butterfly (Troides aeacus) Sheding Forest park, Kenting National Park, Taiwan
Malayan night heron (Gorsachius melanolophus) Taipei city park, Taiwan
Taiwan / Formosan Reevess muntjac baby, (Muntiacus reevesi formosanus), Walami cabin, Yushan National Park, Taiwan. Endemic subspecies
Cloud forest in the Yushan National Park, Taiwan
Fern forest in the Yushan National Park, Taiwan
Black-faced Spoonbill, (Platalea minor), Tainan, Taiwan. Endangered species
Taiwan blue magpie, (Urocissa caerulea), in Taipeh, Taiwan, endemic species
Taiwan blue magpie, (Urocissa caerulea) flying, in Taipeh, Taiwan, endemic species
Taiwan Bar-wing (Actinodura morrisoniana) an endemic species, feeding on red berry in tree, Lu Shan, Taiwan
Taiwan Yellow tit, (Machlolophus holsti) Lu Shan, Taiwan endemic species
Bamboo, Daxueshan National Recreational Forest, Taiwan
Formosan rock macaque (Macaca cyclopis) social grooming behaviour. Alishan National Recreational Forest, Taiwan
Misty mountains, Alishan National Recreational Forest, Taiwan
Taiwan / Formosan yellow-throated marten (Martes flavigula chrysospila) Alishan National Recreational Forest, Taiwan
Swinhoes pheasant (Lophura swinhoii) male, endemic species, Yunlinshan, Taiwan
Black-eared kite, (Milvus migrans lineatus), Bi Tan Bay, Taiwan
Curlew (Numenius arquata) in flight, silhouetted at sunset, with the industrial buildings of Teeside in the background. Durham, UK. January
Canada goose (Branta canadensis) flock taking off from a lake at sunrise. London, UK. November
Pomarine skua (Stercorarius pomarinus) in flight. Durham, UK. September
Glossy ibis (Plegadis falcinellus) in marshland. Donana National Park, Spain
Granitic boulders at Karlu Karlu / Devils Marbles Conservation Reserve, framed by Eucalyptus tree trunks. Northern Territory, Australia. 2008
Woodchat shrike (Lanius senator) perched on bramble bush. Donana National Park, Spain. March
Snowflake moray (Echidna nebulosa) Green Island, Taiwan
Semicircle angelfish (Pomacanthus semicirculatus) Kenting National Park, Kenting, Taiwan
Bargibants seahorse (Hippocampus bargibanti) Kenting Nationalpark, Kenting, Taiwan
Tiny hydrozoa, which is a predatory jellyfish-like animal. The hydrozoans are related to jellyfish and corals and belong to the phylum Cnidaria, Green Island, Taiwan
Juvenile Flying fish (Cypselurus sp), Longdong, Northeast, Taiwan
Peacock butterflies (Aglais io) feeding on Echinops, Norfolk, England, UK, July
Spotted gully shark (Triakis megalopterus) in coral reef. Simons Town, Western Cape, South Africa. June
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Red stingray (Hemitrygon akajei) with Banded houndshark (Triakis scyllium) group and smaller fish in background. Chiba Prefecture, Honshu, Japan. September
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Redspotted catshark (Schroederichthys chilensis) on sea floor amongst Sea urchins (Echinoidea). Las Tacas, Chile. November
Pyjama shark (Poroderma africanum) in reef. Simons Town, Western Cape, South Africa. June
Pyjama shark (Poroderma africanum), two in reef, amongst kelp. Simons Town, Western Cape, South Africa. May
Happy Eddie (Haploblepharus edwardsii) egg case / mermaids purse. False Bay, Western Cape, South Africa. June
Pacific starry skate (Beringraja stellulata) on sea floor. Vancouver Island, British Columbia, Canada. August
Nurse shark (Ginglymostoma cirratum) swimming below surface at sunset. South Bimini Island, Bahamas
Longtail stingray (Hypanus longus) on reef, Whitetip reef shark (Triaenodon obesus) group in background. Socorro Island, Mexico
Leopard houndshark (Triakis semifasciata) group swimming amongst rays of light. San Diego, California, USA
Lemon shark (Negaprion brevirostris) below surface. Tiger Beach, Little Bahama Bank, Bahamas
Eastern shovelnose ray (Aptychotrema rostrata) on sea floor. Nelson Bay, New South Wales, Australia. July
Dusky shark (Carcharhinus obscurus) portrait. Wild Coast, Eastern Cape, South Africa. June
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Circular stingaree (Urolophus circularis) above sea floor. Albany, Western Australia. October
Bronze whaler shark (Carcharhinus brachyurus) group feeding alongside Cape gannet (Morus capensis) during sardine run. Wild Coast, Eastern Cape, South Africa. June
Banded wobbegong shark (Orectolobus halei) resting on reef, Bullseye fish (Pempherididae) shoal in background. South West Rocks, New South Wales, Australia
Banded houndshark (Triakis scyllium) group above sea floor. Chiba Prefecture, Honshu, Japan. October
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
Lion (Panthera leo), females interacting, with cub, Masai Mara, Kenya
Black Woodpecker (Dryocopus martius), male feeding chicks, Bavaria, Germany
Red-backed shrike (Lanius collurio), female with chicks gaping, begging for food, Germany
Spoonbill ( Platalea leucorodia) wading with yellow Water-lily flowers, Hungary
Barbary lion (Panthera leo leo) female, captive, extinct in the wild, occurred in N. Africa
Javan leopard (Panthera pardus melas) captive, endemic to Java. Endemic. Critically endangered species
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Leopard shark (Stegostoma fasciatum) cruising along the edge of an underwater drop off, Philippines