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Group of Clarion angelfish (Holacanthus clarionensis) cleaning an Oceanic manta ray (Mobula birostris) as it swims over the reef, Revillagigedo Islands, Mexico, Pacific Ocean
Two Oceanic manta rays (Mobula birostris) waiting to be cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Silvertip shark (Carcharhinus albimarginatus) portrait, Revillagigedo Islands, Mexico, Pacific Ocean
School of Burrito grunts (Anisotremus interruptus) huddling together in tight ball, Revillagigedo Islands, Mexico, Pacific Ocean
Nudibranch (Phyllidia varicosa) portrait on white background, Bitung, North Sulawesi, Indonesia
Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp)Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp.) on a reef, Ari Atoll, Maldives, Indian Ocean
Great white shark (Carcharodon carcharias) swimming close to the surface, Guadalupe Island, Baja California, Mexico, Pacific Ocean
Sabre squirrelfish (Sargocentron spiniferum) sheltering beneath a large Table coral (Acropora sp.) on a coral reef, Laamu Atoll, Maldives, Indian Ocean
Shoal of large reef fish including Onespot snapper (Lutjanus monostigma), Oriental sweetlips (Plectorhinchus vittatus) and Sabre squirrelfish (Sargocentron spiniferum)
Hawksbill turtle (Eretmochelys imbricata) feeding on a coral reef, Ari Atoll, Maldives, Indian Ocean. Critically endangered
Roundhead parrotfish (Chlorurus strongycephalus) feeding on coral and algae growing on a reef, South Male Atoll, Maldives, Indian Ocean
Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp)Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll, Inner Hebrides, Scotland, UK
Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp)Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp.) on a reef, Similan Islands, Thailand, Andaman Sea
Coral reef below the cliffs of Shark Observatory, Ras Mohammed National Park, Sinai, Egypt, Red Sea
Blue lion's mane jellyfish (Cyanea lamarckii) drifting through a seaweed forest with Red pom-pom seaweedBlue lion's mane jellyfish (Cyanea lamarckii) drifting through a seaweed forest with Red pom-pom seaweed (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll, Inner Hebrides
Bicolor parrotfish (Cetoscarus bicolor) male, scales detail, Sharm El Sheikh, Sinai, Egypt, Red Sea
Two Tawny nurse sharks (Nebrius ferrugineus) on seabed at dusk, Vavuu Atoll, Maldives, Indian Ocean
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Two Dartmoor ponies (Equus ferus caballus), endangered rare breed, colts greeting one another. Dartmoor National Park, Devon, England. June
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
Brent geese (Branta bernicla) small flock in flight over coastal feeding grounds, Lindisfarne National Nature Reserve, Northumberland, UK. February
Leopard (Panthera pardus) hiding in dense foliage, Bardia National Park, Terai, Nepal
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Changeable hawk-eagle / Crested hawk-eagle (Nisaetus cirrhatus) perched on branch, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Asian elephant (Elephas maximus indicus) male, eye detail through foliage, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Grey langurs (Semnopithecus entellus) family group, sitting and climbing in tree, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis) herd in the forest, with just their antlers visible. Bardia National Park, Terai, Nepal
Stork-billed kingfisher (Pelargopsis capensis) perched on branch with beak open, Bardia National Park, Terai, Nepal
Anna's hummingbird (Calypte anna), male flying. Sierra de San Pedro Martir National ParkAnna's hummingbird (Calypte anna), male flying. Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. November
Olive ridley turtle (Lepidochelys olivacea) hatchling, newborn, walking towards Pacific Ocean during arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Red warbler (Cardellina rubra) perched on branch. Milpa Alta forest, outskirts of Mexico City, September
Harlequin Shrimp (Hymenocera elegans) on sea floor. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Misty Pine (Pinus) forest. Milpa Alta forest, outskirts of Mexico City, Mexico. June
Olive ridley turtle (Lepidochelys olivacea) laying eggs in sand under starry night during arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Olive ridley turtle (Lepidochelys olivacea) returning to Pacific ocean after arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Three Scalefin anthias (Pseudanthias squamipinnis) females, joining a larger male on a spawning rise, Sha'ab Abu Nuhas, Egypt, Red Sea
Scalefin anthias (Pseudanthias squamipinnis) male, swimming close to surface, Sharm El Sheikh, Sinai, Egypt, Red Sea
Oceanic manta ray (Mobula birostris) with cephalic fins curled as classic devil ray horns, accompanied by two Remoras (Remora remora), Revillagigedo Islands, Mexico, Pacific Ocean
Dartmoor pony (Equus ferus caballus), endangered rare breed, stallion trotting. Dartmoor National Park, Devon, England. June
Bengal tiger (Panthera tigris tigris) standing in river, whipping water with tail, Bardia National Park, Terai, Nepal. Endangered
Bengal tiger (Panthera tigris tigris) standing on forest path, looking back, Bardia National Park, Terai, Nepal. Endangered
Bengal tiger (Panthera tigris tigris) standing in dense foliage, Bardia National Park, Terai, Nepal. Endangered
Asian elephant (Elephas maximus indicus) male, standing in long grass, Bardia National Park, Terai, Nepal
Two Greater one-horned rhinoceros (Rhinoceros unicornis) walking out of river, Bardia National Park, Terai, Nepal
