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Pyramidal orchid (Anacamptis pyramidalis) in flower, Hampshire, UK June. Focus stacked
Chapuline grasshopper (Sphenarium mexicanum) portrait. Los Tuxtlas rainforest, Mexico, . July
Oceanic manta ray (Mobula birostris) rolling over on its back as it is cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Silver birch (Betula pendula) trees coated in hoar frost in winter, view up into canopy against blue sky, UK. January
Vivid sunset over mountains at Valle de la Luna near Cusi Cusi village, Jujuy, Argentina. October, 2022
Cornflower (Centaurea cyanus) in flower, Surrey, UK. March
Corncockle (Agrostemma githago) and Corn marigolds (Chrysanthemum segetum) flowering in sown wildflower meadow, Surrey, UK. March
Common dodder (Cuscuta epithymum) parasitic on Gorse (Ulex sp. ), Land's End, Cornwall, UK. SeptemberCommon dodder (Cuscuta epithymum) parasitic on Gorse (Ulex sp.), Land's End, Cornwall, UK. September
Ash (Fraxinus excelsior) tree in spring. Surrey, UK. May
Black eyed susan (Rudbeckia hirta) cultivated form, in flower
Common poppy (Papaver rhoeas) flower with bud and seed pod, Surrey, UK. July
Scrapemouth mbuna cichlid (Labeotropheus trewavasae) swimming. Thumbi West island, Lake Malawi, Malawi
Small green utaka cichlid (Nyassachromis leuciscus) swimming. Thumbi West island, Lake Malawi, Malawi
Two Florida manatees (Trichechus manatus latirostris) close to the surface in shallow water, Three Sisters Spring, Crystal River, Florida, USA. Endangered
Two Pantropical spotted dolphins (Stenella attenuata) side by side, porpoising. Both animals have wounds on their bodies inflicted by Cookie cutter shark (Isistius brasiliensis), Hawaii, Pacific Ocean
Tiger shark (Galeocerdo cuvier) swimming over sandy seabed, Tiger Beach. Bahamas, Atlantic Ocean
Sailfin tang (Zebrasoma veliferum) on coral reef at night, Indonesia, Indo-Pacific
Bullseye reef lobster (Hoplometopus holthuisi) on rocky seabed, Hawaii, Pacific Ocean
Crested caracara (Caracara cheriway), adult, walking on beach. Pacific coast, Oaxaca state, Mexico. November
Harlequin Shrimp (Hymenocera elegans) on sea floor. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Orange colored dichromanthus orchid (Dichromanthus aurantiacus) in flower. Milpa Alta forest, outskirts of Mexico City, Mexico. June
Katydid (Copiphora) portrait. Los Tuxtlas Biosphere Reserve, Mexico. JulyKatydid (Copiphora) portrait.Los Tuxtlas Biosphere Reserve, Mexico. July
Red warbler (Cardellina rubra) perched on branch. Milpa Alta forest, outskirts of Mexico City, Mexico. September
Crested caracara (Caracara cheriway) juvenile, on beach. Oaxaca state, Mexico. November
Panamic cushion sea star (Pentaceraster cumingi) detail. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Broadleaf woodsorrel (Oxalis latifolia) in flower, close up. Milpa Alta forest, outskirts of Mexico City, Mexico. August
Mexican violetear hummingbird (Colibri thalassinus) feeding on pink wildflower. Milpa Alta forest, outskirts of Mexico City, Mexico. September
Misty Pine (Pinus) forest. Milpa Alta forest, outskirts of Mexico City, Mexico. June
Olive ridley turtle (Lepidochelys olivacea) newborn hatchling arriving at Pacific ocean during arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Emerald cicada (Zammara smaragdina) on leaves. Montes Azules Biosphere Reserve, Chiapas state, Mexico
Broad-nosed weevil (Exophthalmus opulentus) on leaf. Los Tuxtlas Biosphere Reserve, Mexico
Golden-edged owl-butterfly (Caligo uranus), butterfly captured for the collectors market, wing detail. Montes Azules Biosphere Reserve, Chiapas state, Mexico
Jewel weevil (Eurhinus magnificus) on stem. El Cielo Biosphere Reserve, Mexico
Flat-faced longhorn beetle (Deliathis incana) on branch. Yucatan Peninsula, Quintana Roo state, Mexico
Harlequin beetle (Acrocinus longimanus) headshot close up. Los Tuxtlas Biosphere Reserve, Mexico
Spined orbweaver spider (Micrathena gracilis) on leaf. El Cielo Biosphere Reserve, Mexico
Nudibranch (Goniobranchus geminus) on a coral reef, dorsal view, portrait, Similan Islands, Thailand, Andaman Sea
Longhorn cowfish (Lactoria cornuta) juvenile, swimming over seabed at night, Bitung, North Sulawesi, Indonesia, Molucca Sea
Reef manta ray (Mobula alfredi) feeding on plankton aggregating in the lights from a boat at night, Ari Atoll, Maldives, Indian Ocean
Common clownfish (Amphiprion ocellaris) in the tentacles of its host, Magnificent sea anemone (Heteractis magnifica), on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
School of Scalefin anthias (Pseudanthias squamipinnis) females, swimming around Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents, Ras Mohammed National Park
Nudibranch (Chromodoris annae) on a coral reef, Bitung, North Sulawesi, Indonesia, Molucca Sea
Orange icing sponge (Mycale laevis) growing on Mountainous star coral (Orbicella faveolata) at an early stage of growth, Cayman Islands, Caribbean Sea
Pair of Spotted unicornfish (Naso brevirostris) courtship display above the reef, Ari Atoll, Maldives, Indian Ocean
Scalefin anthias (Pseudanthias squamipinnis) male, swimming close to surface, Sharm El Sheikh, Sinai, Egypt, Red Sea
Pair of Nudibranchs (Flabellina exoptata) just prior to mating, the mating apparatus of the one on left is visible, Bitung, North Sulawesi, Indonesia, Molucca Sea
Group of Clarion angelfish (Holacanthus clarionensis) cleaning an Oceanic manta ray (Mobula birostris) as it swims over the reef, Revillagigedo Islands, Mexico, Pacific Ocean
Two Oceanic manta rays (Mobula birostris) waiting to be cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Silvertip shark (Carcharhinus albimarginatus) portrait, Revillagigedo Islands, Mexico, Pacific Ocean
School of Burrito grunts (Anisotremus interruptus) huddling together in tight ball, Revillagigedo Islands, Mexico, Pacific Ocean
Nudibranch (Phyllidia varicosa) portrait on white background, Bitung, North Sulawesi, Indonesia
Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp)Yellowmargin moray (Gymnothorax flavimarginatus) peering out from Digitate leather coral (Sinularia sp.) on a reef, Ari Atoll, Maldives, Indian Ocean
Great white shark (Carcharodon carcharias) swimming close to the surface, Guadalupe Island, Baja California, Mexico, Pacific Ocean
Sabre squirrelfish (Sargocentron spiniferum) sheltering beneath a large Table coral (Acropora sp.) on a coral reef, Laamu Atoll, Maldives, Indian Ocean
Shoal of large reef fish including Onespot snapper (Lutjanus monostigma), Oriental sweetlips (Plectorhinchus vittatus) and Sabre squirrelfish (Sargocentron spiniferum)
Hawksbill turtle (Eretmochelys imbricata) feeding on a coral reef, Ari Atoll, Maldives, Indian Ocean. Critically endangered
Roundhead parrotfish (Chlorurus strongycephalus) feeding on coral and algae growing on a reef, South Male Atoll, Maldives, Indian Ocean
Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp)Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll, Inner Hebrides, Scotland, UK
Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp)Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp.) on a reef, Similan Islands, Thailand, Andaman Sea
Coral reef below the cliffs of Shark Observatory, Ras Mohammed National Park, Sinai, Egypt, Red Sea
Blue lion's mane jellyfish (Cyanea lamarckii) drifting through a seaweed forest with Red pom-pom seaweedBlue lion's mane jellyfish (Cyanea lamarckii) drifting through a seaweed forest with Red pom-pom seaweed (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll, Inner Hebrides
Bicolor parrotfish (Cetoscarus bicolor) male, scales detail, Sharm El Sheikh, Sinai, Egypt, Red Sea
Two Tawny nurse sharks (Nebrius ferrugineus) on seabed at dusk, Vavuu Atoll, Maldives, Indian Ocean
Warty frogfish (Antennarius maculatus) yawning in its hiding place amongst white sponges, Bitung, North Sulawesi, Indonesia, Molucca Sea
Two Dartmoor ponies (Equus ferus caballus), endangered rare breed, colts greeting one another. Dartmoor National Park, Devon, England. June
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
Brent geese (Branta bernicla) small flock in flight over coastal feeding grounds, Lindisfarne National Nature Reserve, Northumberland, UK. February
Leopard (Panthera pardus) hiding in dense foliage, Bardia National Park, Terai, Nepal
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Changeable hawk-eagle / Crested hawk-eagle (Nisaetus cirrhatus) perched on branch, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Asian elephant (Elephas maximus indicus) male, eye detail through foliage, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Grey langurs (Semnopithecus entellus) family group, sitting and climbing in tree, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis) herd in the forest, with just their antlers visible. Bardia National Park, Terai, Nepal
Stork-billed kingfisher (Pelargopsis capensis) perched on branch with beak open, Bardia National Park, Terai, Nepal
Anna's hummingbird (Calypte anna), male flying. Sierra de San Pedro Martir National ParkAnna's hummingbird (Calypte anna), male flying. Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. November
Olive ridley turtle (Lepidochelys olivacea) hatchling, newborn, walking towards Pacific Ocean during arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Red warbler (Cardellina rubra) perched on branch. Milpa Alta forest, outskirts of Mexico City, September