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Hooded grebes (Podiceps gallardoi) pair dancing in breeding display, on a lake above the Buenos Aires plateau, Patagonia National Park, Santa Cruz, Argentina. Cropped
Ocelot (Leopardus pardalis) climbing a tree trunk Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Lightning strike over South Africa's largest coal processing plant. Sasol, Mpumalanga, South Africa, November, 2016Lightning strike over South Africa's largest coal processing plant. Sasol, Mpumalanga, South Africa, November, 2016
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Female Black-footed cat (Felis nigripes) with kitten, at edge of burrow. These animals have been radio collared as part of the Black-footed Cat Project, Karoo South Africa
Female Black-footed cat (Felis nigripes) in the desert at night, Karoo, South Africa. Taken on location for the BBC series Big Cats
Mexican fiddler crab (Leptuca crenulata) small male displaying with larger individual behind, Morua Estuary, near Rocky Point (Puerto Penasco), Gulf of California, Mexico, June
Wild Indian Ocean bottlenose dolphin (Tursiops adunctus) swimming with a golden retriever dog (Canis lupus familiaris). Gubal Island, Egypt, Red Sea, Indian Ocean
Mute swan (Cygnus olor) landing on frozen lake, Richmond Park, London, UK. February
Mute swan (Cygnus olor) swimming in a small pool of frozen lake, Richmond Park, London, UK. February
Female Fox (Vulpes vulpes) caught in a shaft of light on the porch of a farmhouse, Verteskozma, Hungary
Fox (Vulpes vulpes) drinking water from a sauna pool in a garden; low angle view looking up through water, Verteskozma, Hungary
Female Fox (Vulpes vulpes) walking on tree trunk in heavy snowstorm in the dark, Verteskozma, Hungary
Female Red fox (Vulpes vulpes) foraging inside a rotting tree trunk, Verteskozma, Hungary. Hungary Nature Photographer of the Year Competition 2021 Highly Commended - Mammals Behaviour category
African elephant (Loxodonta africana) hidden behind a bush, Marataba Private Reserve, Marakele National Park, South Africa
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
Male Bleekers parrotfish (Chlorurus bleekeri) races across a coral reef as it displays to females.Yanggefo Island, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Weedy seadragon (Phyllopteryx taeniolatus) swimming over seaweeds. Sydney, New South Wales, Australia. Pacific Ocean
Crescent-tail bigeye (Priacanthus hamrur) school on a coral reef, shot with a long exposure. North Male Atoll, Maldives. Indian Ocean
Male Fifteen spined stickleback / Sea stickleback (Spinachia spinachia) guarding a nest in Spaghetti seaweed / Thong weed (Himanthalia elongata), Isle of Coll, Inner Hebrides, Scotland, UK
Young, female Grey seal (Halichoerus grypus) swimming over a forest of Cuvie kelp (Laminaria hyperborea), Inner Hebrides, Scotland, UK, Atlantic Ocean. August
A tangle of Spaghetti seaweed / Thong weed (Himanthalia elongata) growing beneath the straight basalt columns outside Fingals Cave. Isle of Staffa, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Common dolphin (Delphinus delphis) porpoising at the surface to breathe, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Adult Shags (Phalacrocorax aristotelis) with young, gathered on rocks, Treshnish Isles, Southern Hebrides, Scotland, UK, Atlantic Ocean. August
Common dolphin (Delphinus delphis) leaping out of sea, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Snakelocks anemone (Anemonia viridis) growing on a stalk of Eelgrass (Zostera marina) on the seabed, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Spaghetti seaweed / Thong weed (Himanthalia elongata) forest reaching up to water surface to absorb sunlight, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Compass jellyfish (Chrysaora hysoscella) swimming near surface, Cornwall, UK, English Channel
Common starfish (Asterias rubens) with pink encrusting algae, Farne Islands, Northumberland, UK, North Sea
Aggregation of Common sea urchins (Echinus esculentus) beneath Cuvie kelp (Laminaria hyperborea) forest, Farne Islands, Northumberland, UK, North Sea
Detail of Light-bulb sea squirts (Clavelina lepadiformis), Swanage, Dorset, UK, English Channel
Leachs spider crab (Inachus phalangium) sheltering beneath the stinging tentacles of Snakelocks anemone (Anemonia viridis). Swanage, Dorset, UK, English Channel
Small Stalked jellyfish (Calvadosia sp. ) attached to Bootlace weed / MermaidSmall Stalked jellyfish (Calvadosia sp.) attached to Bootlace weed / Mermaids hair (Chorda filum), Pendennis Head, Cornwall, UK. English Channel
A large herbivorous Seahare (Aplysia punctata) crawling around a Common eelgrass (Zostera marina) meadow, Helford River Estuary, Cornwall, UK
Grey top-shell (Gibbula cineraria) grazing on Golden kelp (Laminaria ochroleuca). Pendennis Head, Cornwall, UK, English Channel
Young Long clawed squat lobster (Munida rugosa), portrait, Loch Linnhe, The Highlands, Scotland, UK
Spider crab (Hyas sp. ) scaling the stipe of Cuvie kelp (Laminaria hyperborea)Spider crab (Hyas sp.) scaling the stipe of Cuvie kelp (Laminaria hyperborea) up into the canopy of the kelp forest. Loch Carron, Highlands, Scotland, UK
Common hermit crab (Pagurus bernhardus) feeding on a Flame shell (Limaria hians) in a kelp (Laminaria hyperborea) forest, Loch Carron, Highlands, Scotland, UK
Nudibranch (Fjordia lineata) crawling amongst the legs of a Black brittlestar (Ophiocomina nigra), Loch Carron, Highlands, Scotland, UK
Male Lumpsucker (Cyclopterus lumpus) swimming through Kelp forest (Laminaria hyperborea), Loch Carron, Highlands, Scotland, UK
Pair of Spot gobies (Gobiusculus flavescens) nesting in a old Common sea urchin (Echinus esculentus) shell, Loch Carron, Highlands, Scotland, UK
Face of an Anglerfish (Lophius piscatorius), Chesil Beach, Dorset, UK, English Channel
Detail of a St Johns stalked jellyfish (Calvadosia cruxmelitensis) growing on seaweed, Chesil Beach, Dorset, UK, English Channel
RF - Tompot blenny fish (Parablennius gattorugine) peering out from hole under Swanage Pier, Dorset, UK, (This image may be licensed either as rights managed or royalty free.)
Hooded grebe (Podiceps gallardoi) pair on lake with chick perched on one parents back, second parent feeding chick with invertebrate, Strobel plateau, Santa Cruz, Patagonia, Argentina. Cropped
Hooded grebe (Podiceps gallardoi) on lake, Strobel plateau, Santa Cruz, Patagonia, Argentina
Hooded grebes (Podiceps gallardoi) pair on lake, with mountains in Chile in background, Buenos Aires plateau, Patagonia National Park, Santa Cruz, Argentina. January 2018
Greater hoopoe-lark (Alaemon alaudipes) standing on desert sand, Southern Morocco, Africa. Cropped
Erg Chebbi sand dunes, Sahara desert, Southern Morocco, Africa. January, 2020
Guelta / water ponds in the low Draa valley. Southern Morocco, Africa. March, 2020
Painted lady (Vanessa cardui) sucking nectar from an acacia flower during spring migration. Aydar, Southern Morocco, Africa
Desert sparrow (Passer simplex) perched in long grass, Aousserd, Western Sahara, Africa
Small-scale cultivation, Ramlia Oasis, Sahara desert, Southern Morocco, Africa
Rhinoceros rock art, Sahara desert, Ait Ouaazik, Southern Morocco, Africa
Sahara desert landscape taken from Erg Ouzina. Southern Morocco, Africa. December, 2009
Black stork (Ciconia nigra) in flight during spring migration across the Sahara desert, Aydar, Southern Morocco, Africa
Lesser Egyptian jerboa (Jaculus jaculus) in the desert at night, Oued Afra, Western Sahara, Africa
Female Cuviers gazelle (Gazella cuvieri) calf, age 6 months, with its mother, Djebel Ouarkziz, Southern Morocco, Africa. Customized camera-trap
Desert monitor (Varanus griseus) on rocky desert landscape, Aydar, Southern Morocco, Africa
Acacia tree at dusk, Sahara desert, Djebel Ouarkziz, Southern Morocco, Africa. March, 2016
Sand dunes below the northern slopes of Djebel Ouarkziz, Sahara desert, Southern Morocco, Africa. February, 2020
Landscape showing diversity of Saharan ecosystem, Sahara desert, Oued Ziz from Erg Ouzina, Southern Morocco, Africa. December, 2009
Desert vegetation in the dunes, Sahara desert, Erg Chebbi, Southern Morocco, Africa. January, 2010
Rain and storm clouds passing over the desert, Karoo, South Africa, 2016
Fishing cat (Prionailurus viverrinus) with two kittens, age 4 weeks, learning to hunt for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss
Fishing cat (Prionailurus viverrinus) with kitten, age 4 weeks, in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss, NW Bangladesh 2017
Fishing cat (Prionailurus viverrinus) hunting for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss, NW Bangladesh 2017
Female Fishing cat (Prionailurus viverrinus) hunting for fish in wetlands, photographed during a release project to relocate Fishing cats affected by habitat loss, NW Bangladesh 2017
Male Jaguar (Panthera onca) walking through rainforest, Belize, Central America, 2017. Captive. Filmed for the BBC series Big Cats
Pair of Leopard tortoises (Stigmochelys pardalis) mating, Karoo, South Africa
Margay (Leopardus wiedii) on tree branch, Belize, Central America. 2017. Captive
Smoke billowing from Mount Bromo volcano. View from the summit of Mount Batok. Bromo Tengger Semeru National Park, Java, Indonesia, July, 2013
Ocelot (Leopardus pardalis) sitting on forest floor, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) in rainforest, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) hiding amongst tree roots, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) looking up, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Ocelot (Leopardus pardalis) high up in tree, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Pallas's cat (Otocolobus manul) kitten waiting at den for mother to return, Mongolia. June. 2017Pallas's cat (Otocolobus manul) kitten waiting at den for mother to return, Mongolia. June. 2017. Filmed for BBC series Big Cats
Female Pallass cat (Otocolobus manul) suckling four young kittens at their den, Mongolia, June, 2017. Filmed for BBC series Big Cats
Large flower of the parasitic plant Rafflesia pricei, growing in rainforest, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Hoffmanns two-toed sloth (Choloepus hoffmanni) hanging from tree branch, Panama, Central America
Male Bengal tiger (Panthera tigris tigris) walking through mangrove forest, Sundarbans, India, 2017