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Yew berries (Taxus baccata) Berwickshire, Scotland, September
Highland Cattle (Bos tarus) herd grazing, West Highland Way, Tyndrum, Scotland, UK, August
Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica
View up into Beech tree (Fagus sylvatica) in autumn, Berwickshire, Scotland, UK, October
Fly Agaric Fungus (Amanita muscaria) within forestry plantation, Strathspey, Scotland, UK, September
Curlew (Numenius arquata) roosting at high tide amongst Oystercatchers {Haematopus ostralegus} Northumberland, England, September
Female River otter (Lutra lutra) on sea loch, Torridon, Wester Ross, Scotland, July
African lion (Panthera leo) male in habitat, Okavango Delta, Botswana, January
Dew on cobweb, Berwickshire, Scotland, UK, September
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA
African elephant (Loxodonta africana) juvenile flapping ears, Okavango Delta, Botswana
Long exposure of wave breaking on rocks at sunset, Caithness, Scotland, August
Wych Elm (Ulmus glabra) dead trees in mist at dawn, Berwickshire, Scotland, July
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring, Crystal River, Florida, USA
Golden Eagle (Aquila chrysaetos) controlled, close-up of falconers bird, Southern Scotland, UK, February
Badger (Meles meles) cub inside hollow log, Berwickshire, Scotland, UK, June
Norway Maple (Acer platanoides) fallen leaves in autumn, Queen Elizabeth Forest Park, Stirlingshire, Scotland, October
Southern stingray (Hypanus americanus) swimming over sand ripples on sandbar, Stingray City, Grand Cayman. British West Indies
African elephant (Loxodonta africana) juvenile holding up trunk, Okavango Delta, Botswana
Silhouette of spider at sunset, Berwickshire, Scotland, August
Quiraing, Trotternish Ridge, Isle of Skye, Scotland, December
Pinnacles of Old Man of Stoer, Isle of Skye, Scotland, June
Bluebells (Hyacinthoides non-scripta) flowering in oakwood, Perthshire, Scotland, UK, May
King Penguins (Aptenodytes patagonicus) head portraits, on the beach at Salisbury Plain, South Georgia, South Atlantic. January
Northern gannet (Morus bassanus) gathering nesting material, Bass Rock, Firth of Forth, East Lothian, UK, July
King Penguin (Aptenodytes patagonicus) displaying on the beach front. Salisbury Plain, South Georgia, Antarctica. January, sequence 5/6
King Penguin (Aptenodytes patagonicus) displaying on the beach front. Salisbury Plain, South Georgia, Antarctica. January, sequence 4/6
Five King Penguins (Aptenodytes patagonicus) standing together on the beach at Salisbury Plain, South Georgia, South Atlantic. January