Heather Angel Collection

Cedar (Cedrus sp), male catkins releasing pollen. Surrey England, UK

Hong Kong orchid tree (Bauhinia blakeana) flowers. Hong Kong, December

Giant panda (Ailuropoda melanoleuca) sliding down slope on back after losing footing. Panda Breeding Centre, Wolong, Sichuan, China. January 1987

Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July

Wild sage (Lantana camara), flowers turn pink following pollination. Native to tropical America. Invasive species, Tanzania

Pygmy Rwandan water lily (Nymphaea thermarum) in cultivation at Kew Gardens, London, England, UK. Extinct in the wild. Native to thermal springs, Rwanda

Mysore clock vine (Thunbergia mysorensis) with nectar overflowing in morning. In cultivation, Kew Gardens, London, England, UK. Native to India

Wood forget-me-not (Myosotis sylvatica) flowers, yellow corollas on fresher flowers. In damp woodland, Surrey, England, UK. May

Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation

Black coral vine (Kennedia nigricans) flowers, pollinated by honeyeaters (flowers triggered by honeyeater, releasing the stamens, style and stigma), Western Australia Sequence 2 of 2

Stigmas in female flowers of single Wax begonia (Begonia semperflorens group) mimic pollen laden stamens produced by male flowers but offer no reward to pollinators

Seep monkey flower (Mimulus guttatus), bifid stigma above stamens. Nectar spot guides and central groove leading pollinators to nectar. Focus stacked

Prickly comfrey (Symphytum asperum), flowers open red and change to blue. Caucasus, Russia. June

Prophet flowers (Arnebia pulchra) with dark spots that act as nectar guides. Caucasus, Russia. June

Common rue (Ruta graveolens) flower. Cultivated in herb garden, Surrey, England, UK. Native to Balkan Peninsula

Voodoo lily (Amorphophallus bulbifer). Native to Asia

Fairy fern (Azolla filiculoides) and Water forget me not (Myosotis scorpioides) in frozen garden pond. Surrey, England, UK. January

St Helena ebony / Dwarf ebony (Trochetiopsis ebenus) with secondary pollen presentation on petal rims. In cultivation, Kew Gardens, London, England, UK. Endemic to St Helena, South Atlantic Ocean

Chilean bellflower (Crinodendron hookerianum) releasing pollen via sonication which replicates buzz pollination. Native to Chile. Controlled conditions

Congo cockatoo (Impatiens niamniamensis) flower, backlit with nectar level visible in spur. Cultivated in glasshouse. Native to Africa

Crown imperial lily (Fritillaria imperialis), six nectaries at base of stamens and stigma. In cultivation, Native to southeast Turkey and western Himalayas

Darwins slipper flower (Calceolaria uniflora) in UV light. Stamens and fleshy lip that provides food reward for polliator Least seedsnipe (Thinocorus rumicivorus) fluorescing

Morning glory (Ipomoea tricolor) flower with pollen grains scattered by visiting insects

Celandine crocus (Crocus korolkowii) carpet in area of snowmelt. Pollinated by solitary bees and bumblebees. In valley, near Ansob Pass, Tajikistan. April 2014

Musquia (Musschia aurea) flower. Cultivated in glasshouse, UK. Pollinated by Lizard (Lacerta dugesii) and Madeira bumblebee (Bombus maderensis) in Madeira

Jade vine (Strongylodon macrobotrys) in glasshouse at Kew Gardens, London, UK. Endemic to the Philippines. Pollinated by sunbirds

Winter aconite (Eranthis hyemalis) with ring of nectaries below stamens. Focus stacked

Foxglove (Digitalis purpurea), blue nectar guides and anthers, pollen and stigma fluorescing in UV light. Focus stacked

Skunk cabbage (Lysichiton americanus) in visible light. In cultivation, Surrey, England, UK. Native to Canada and USA. Sequence 1/2

Large bee-fly (Bombylius major) nectaring on Lungwort (Pulmonaria sp). In garden, Surrey, England, UK. March

Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China

Regal lily (Lilium regale) with Crab spider (Thomisidae sp) waiting to predate insects. Near Maoxian, Min Valley, Sichuan Province, China

Shrubs, trees and lianas in tropical rainforest. Xishuangbanna, Yunnan Province, China

Euphorbia (Euphorbia jolkinii) leaves turning red in autumn. Napa Hai, Shangri-la, Yunnan Province, China. September

Cut bamboo stems stacked in a pile, end on. Anhui Province, China

Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China

Chinese grass carp / White amur (Ctenopharyngodon idella), farmed for food in China. Captive specimen, China

Sacred lotus (Nelumbo nucifera) flowering in Puzhehai Lake with peaks in background. Puzhehei National Wetland Park, Yunnan Province, China. 2009

Mauritian bloody bellflower (Nesocodon mauritianus), female phase with red nectar at base of style. Cultivated at Kew Gardens, London, England, UK. Native to Mauritius

Cats whiskers / Black bat flower (Tacca chantrieri). Pollinated by flies. Cultivated in glasshouse, Surrey, England, UK. Native to Southeast Asia

Butterflies and day flying moths feed from Loofah flower (Luffa cylindrica) and pick up pollen on their legs, underside of thorax and antennae, South Sichuan Bamboo Sea, China

Grand-flowered horned poppies (Glaucium grandiflorum) in southern Turkey, June

Pollen release from cyclamen flower via sonication, which mimics buzz pollination. The structure of the cyclamen flower prevents the pollen getting wet

Day gecko (Phelsuma sp. ) with extended tongue lapping up nectar from crown of thorns
Day gecko (Phelsuma sp.) with extended tongue lapping up nectar from crown of thorns (Euphorbia milii) flower, Madagascar

Chinese peacock butterfly (Papilio bianor) drinking nectar from a Hibiscus flower

Wasp walking around a Spanish love-in-a-mist (Nigella hispanica) flower to sip nectar from ring of blue nectaries

Honey bee (Apis mellifera) with full pollen baskets feeding on Balkan anemone (Anemone blanda). In garden. Surrey, England, UK. April

Spotted blister beetle (Ceroctis capensis) feeding on Gladiolus (Gladiolus alatus) petals. Cape Town, South Africa. September

Honey bee (Apis mellifera) extending proboscis to nectar on Common rue (Ruta graveolens). Green aphid (Aphidoidea) beneath flower. Surrey, England, UK. July

Parasitic wasp (Gasteruption assectator) nectaring on Common rue (Ruta graveolens). Surrey, England, UK. July

Cambridge vagrant butterfly (Nepheronia thalassina) nectaring on Lantana (Lantana camara). Flowers turn from yellow to pink when no nectar is provided. In garden, Tanzania. Native to tropical Americas

Solitary bee (Apoidea) feeding on Crocus (Crocus korolkowii), covered in pollen. Ansob Pass, Tajikistan. April

Honey bee (Apis mellifera) nectaring on freshly opened yellow Lantana (Lantana camara) flowers. Flowers turn pink when nectar no longer provided. In garden, Tanzania. Native to tropical Americas

Bumblebee (Bombus sp) queen feeding on Crocus (Crocus korolkowii), covered in pollen. Ansob Pass, Tajikistan. April

Tree bumblebee (Bombus hypnorum) nectaring on Camellia St Ewe (Camellia u williamsii)
Tree bumblebee (Bombus hypnorum) nectaring on Camellia St Ewe (Camellia × williamsii). Surrey, England, UK. March

Angle shades moth (Phlogophora meticulosa) nectaring on Ivy (Hedera helix) at night. Surrey, England, UK. October

Ivy bee (Colletes hederae) female collecting pollen from Ivy (Hedera helix). Surrey, England, UK. October

Wasp (Hymenoptera) robbing nectar from Henrys lily (Lilium henryi). Surrey, England, UK. August

Honey bee (Apis mellifera) foraging on Peony (Paeonia officinalis) pollen. Surrey, England, UK. May

Bumblebee (Bombus sp) looking for pollen amongst fallen Southern magnolia (Magnolia grandiflora) stamens. Surrey, England, UK. July

Large bee fly (Bombylius major) feeding on Primrose (Primula vulgaris). England, UK. March

Red admiral (Vanessa atalanta) nectaring on Ivy (Hedera helix). September

Honey bee (Apis mellifera) nectaring on Bonnet bellflower (Codonopsis clematidea). Surrey, England, UK. July

Bee-fly (Bombylius sp) nectaring on Rock soapwort (Saponaria ocymoides), blue pollen on head. Alpes Maritimes, France. May

Marmalade hoverflies (Episyrphus balteatus), two feeding on pollen of Dwarf formosa lily (Lilium formosanum var. pricei). Surrey, England, UK. July

Leopard lacewing (Cethosia cyane) nectaring on Tropical milkweed / Bloodflower (Asclepias curassavica). Yunnan, China

Moth (Lepidoptera) nectaring on Bokkeveld evening lily (Hesperantha cucullata) at dusk. Nieuwoudtville, South Africa. August

Sea cabbage (Senecio candicans) amongst pebbles on sandy seashore. Sea Lion island, Falkland Islands

Painted lady butterfly (Vanessa cardui) nectaring on Witch hazel (Hamamelis x intermedia Jelena ). Surrey, England, UK. February

Hibiscus (Hibiscus sp) flower with stigmas and pollen covered stamens. Focus stacked

Orange bush monkeyflower (Mimulus aurantiacus). Native to California and Oregon, USA. Focus stacked

Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removed
Snowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked

Crocus (Crocus sp) flower from above, closing in evening. Focus stacked

Water tulip (Cypella aquatilis) from above, in garden pond. Native to Brazil

Daffodil (Narcissus sp) opening, view from above. Focus stacked

Peach leaved bellflower (Campanula persicifolia) dissection, petals removed to show how stamens transfer pollen to style via secondary pollen presentation in bud stage. Focus stacked

Grand balsam (Impatiens grandis) with long spur. Pollinated by butterflies, native to India and Sri Lanka. Focus stacked, cultivated plant

Figwort (Scrophularia grandiflora) against sky. Globose flower with protruding upper lip that functions like umbrella to protect stamens. Cultivated in garden

Carpenter bee (Xylocopa sp) nectaring on Clary sage (Salvia sclarea). Pollen deposited on bees back by hanging anthers. Turkey. June

Himalayan balsam (Impatiens glandulifera), dissection of flowers. Male phase above, downward projecting stamens release pollen onto backs of bumblebees and honey bees

Rhododendron (Rhododendron sp) flower, anthers and red stigma visible. Garden in Surrey, England, UK

Sweet breath of spring (Lonicera fragrantissima) in garden, Surrey, England, UK. Native to China

Alpine rose (Rhododendron ferrugineum) in montane grassland above treeline. Giau Pass / Passo di giau, Dolomites, Italy. July 2011

Caucasian lily (Lilium monadelphum), Caucasus, Russia. June

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