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Peregrine falcon (Falco peregrinus) flying past crane, Barcelona, Spain, April
Greater black backed gull (Larus marinus) swallowing large fish, North Atlantic, Flatanger, Nord-Trndelag, Norway, August 2008
Golden eagle (Aquila chrysaetos) in tree with dead bird, wings out stretched, Flatanger, Norway, November 2008
Grey heron (Ardea cinerea) with wings out stretched, Elbe Biosphere Reserve, Lower Saxony, Germany, September 2008
Rear view of Black stork (Ciconia nigra) with wings outstretched, Elbe Biosphere Reserve, Lower Saxony, Germany, August 2008
Grey heron (Ardea cinerea) taking off, Texel, Netherlands, May 2009
Plume moth (Pterophoridae family) Fliess, Naturpark Kaunergrat, Tirol, Austria, July 2008 WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Canada goose (Branta canadensis) chasing off urban Red fox (Vulpes vulpes) London, May
Osprey (Pandion haliaetus) in flight carrying fish, Kangasala, Finland, August 2009
Griffon vulture (Gyps fulvus) running with wings stretched, Andorra, June 2009. WWE INDOOR EXHIBITION
Lammergeier (Gypaetus barbatus) chasing away Carrion crow (Corvus corone corone) Cebollar, Torla, Aragon, Spain, November 2008
Two Lammergeiers (Gypaetus barbatus) fighting, Serra de Beumort, Gerri de la Sal, Catalonia, Spain, November 2008. WWE BOOK. WWE OUTDOOR EXHIBITION
Redshank (Tringa totanus) calling in flight, Outer Hebrides, Scotland, UK, June
Female Goosander (Mergus merganser) drying feathers, Gwynedd, Wales, UK, January
An adult male Mallard (Anas platyrhynchos) comes in to land, backlit by evening sunlight, Derbyshire, England, UK, March. Did you know
Adult Coot (Fulica atra) running on the surface of a lake, Derbyshire, England, UK, March 2010. Did you know? Coots can dive up to 2 metres
Eurasian dotterel (Charadrius morinellus) pair mating in breeding habitat on upland plateau of Grampian mountains, Cairngorms NP, Scotland, UK, May 2011
Common Swallowtail Butterfly (Papilio machaon) resting on Vipers Bugloss / Blueweed (Echium vulgare) in alpine meadow. Nordtirol, Tirol, Austrian Alps, Austria, 1700 metres altitude, July
Common blue butterfly (Polyommatus icarus), resting on flowering heather, Arne RSPB reserve, Dorset, England, UK September
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, originating from North and Central America, sequence 2 of 2. Captive
Greater flamingos (Phoenicopterus roseus) mating, in lagoon, Pont Du Gau, Camargue, France, April 2009
Dalmatian pelican (Pelecanus crispus) profile in flight, Danube delta rewilding area, Romania
Bullers shearwater (Puffinus bulleri) in flight low over the water, showing upperwing. Off Whitianga, Coromandel Peninsula, New Zealand
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Common buckeye butterfly (Junonia coenia) feeding on Ironweed (Vernonia altissima) flower, North Guilford, Connecticut, USA, August
Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
Great Tit (Parus major) in flight. Bavaria, Germany, December
Male Comet / Madagascar moon moth (Argema mittrei) recently emerged drying its wings. Andasibe-Mantadia National Park, Madagascar
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Goshawk (Accipiter gentilis) in flight over woodland clearing, Cairngorms National Park, Scotland, UK
Ferruginous hawk (Buteo regalis) landing. Captive
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
RF - Banded darter (Sympetrum pedemontanum) group of five resting, Netherlands, September
Blue-chested hummingbird (Amazilia amabilis) male feeding from flower, Costa Rica, Central America
Coppery-headed Emerald, Elvira cupreiceps, male in flight feeding on Spiral Ginger (Costus Pulverulentus), Central Valley, Costa Rica, Central America, December 2006
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Puffin (Fratercula arctica) landing in snow, Norway, March
Male Keeled skimmer (Orthetrum coerulescens) in flight, Holt Lows CP, Norfolk, England, August
Orange Tip Butterfly (Anthocharis cardamines) female resting on bluebell, Lanhydrock woodland, Cornwall. May 2012
Close-up of Swallowtail (Papilo machaon) wings, Emilia Romagna Region, Italy
Egyptian vulture (Neophron percnopterus) landing, Faia Brava Reserve, Coa valley, Portugal, May
White tailed sea eagle (Haliaetus albicilla) in flight, Flatanger, Norway, June 2008 DUPE OF 1213183; TO DELETE
Scalloped oak moth (Crocallis elinguaria) on twig, Uplyme, Devon, England, July
Jay {Garrulus glandarius} close up of wing feathers Europe
Female Black veined white butterfly (Aporia crataegi) laying eggs on Buckthorn, Lagadin region, Lake Ohrid, Galicica National Park, Macedonia, June 2009
Female Gyrfalcon (Falco rusticolus) landing on rock, Myvatn, Thingeyjarsyslur, Iceland, May 2009
Long tailed skua (Stercorarius longicaudus) in flight, Thingeyjarsyslur, Iceland, June 2009