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Lammergeier / Bearded Vulture (Gypaetus barbatus) in flight. Ordesa National Park, Spanish Pyrenees, April
Siskins (Carduelis spinus) fighting over food. Oisterwijk, The Netherlands, March
Bullers shearwater (Puffinus bulleri) in flight low over the water, showing upperwing. Off Whitianga, Coromandel Peninsula, New Zealand
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
Great Tit (Parus major) in flight. Bavaria, Germany, December
Three Brown-hooded Parrots (Pyrilia haematotis) one taking off from branch, in the rain Laguna del Lagarto, Santa Rita, Costa Rica
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
Male Comet / Madagascar moon moth (Argema mittrei) recently emerged drying its wings. Andasibe-Mantadia National Park, Madagascar
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Greater flamingo (Phoenicopterus roseus) flock standing in lake as others fly overhead and one bird flaps its wings, Camargue, France, January
Blue and yellow macaw (Ara ararauna) in flight, Pantanal, Mato Grosso do Sul, Brazil
Peacock katydid (Pterochroza ocellata) in defensive posture, Yasuni National Park, Orellana, Ecuador
Giant hummingbird (Patagona gigas) in flight, portrait, Cotopaxi National Park, Cotopaxi, Ecuador
Little egret (Egretta garzetta) landing on water, Ubrique, Southern Spain. June
Long-tailed tit (Aegithalos caudatus) feeding on icicle formed from sap of Painted maple tree (Acer pictum). Hokkaido, Japan. February
Atlantic puffin (Fratercula arctica) standing on rock with fish in beak, with wings spread, Machias Seal Island, Maine, USA. July
Great crested grebe (Podiceps cristatus) shaking chick from its back on lake, Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Hairy dragonfly (Brachytron pratense) male, resting on plant stem, Brackagh Moss National Nature Reserve, County Armagh, Northern Ireland, UK. June
Great white egret / Common egret (Ardea alba) preening feathers in shallow water of pond, Baie de Somme Nature Reserve, Hauts-de-France, France. September
Red grouse (Lagopus lagopus) in flight over moorland, Yorkshire, UK. September
Snowy owl (Bubo scandiaca) female, in flight in winter, Canada. January
White-tailed eagle (Haliaeetus albicilla) taking off from iceberg at sunrise, Shiretoko National Park, Hokkaido, Japan
Rhinoceros hornbill (Buceros rhinoceros) male inflight, carrying mouse prey back to nest, Budo-Sungai Padi National Park, Thailand
Wandering albatross (Diomedea exulans) male, standing with wings spread in courtship display, Prion Island, South Georgia
Magnificent riflebird (Ptiloris magnificus alberti) male, courtship display, Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
Goldcrest (Regulus regulus) in flight, portrait, Parainen Uto, Finland. September
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Peacock katydid (Pterochroza ocellata) on leaf in defensive posture, Yasuni National Park, Amazon Basin, Ecuador
Two Canada geese (Branta canadensis) in flight with autumn foliage behind, Maryland, USA. October
Red grouse (Lagopus lagopus) coming in to land on flowering Heather (Calluna vulgaris), Yorkshire, UK. September
Swallowtail butterfly (Papilio machaon) in flight, Finland, June
Welcome swallow (Hirundo neoxena) in flight over pond. Christchurch, New Zealand
Juvenile Kea (Nestor notabilis) in flight. Arthur's Pass National Park, South Island, New Zealand. Endangered Species
Redstart (Phoenicurus phoenicurus) juvenile male, in flight, chasing an insect, Parainen Uto, Finland. September
Nutcracker (Nucifraga caryocatactes) in flight with wings spread, Sipoo, Finland. January
Wood sandpiper (Tringa glareola) in flight, Parainen Uto, Finland. August
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Two Buzzards (Buteo buteo) fighting over pheasant carcass, Marlborough Downs, Wiltshire, UK. December
Male Eastern bluebird (Sialia sialis) feeding on snow-covered Holly (Ilex sp) berries, near Ithaca, New York, USA. January
Male Giant kingfisher (Megaceryle maxima) diving, Allahein River, The Gambia
Common greenshank (Tringa nebularia) wading in river with wings spread, Allahein River, The Gambia
Grey-headed flying-fox (Pteropus poliocephalus) in flight, with tongue out after licking water from its body. Having just completed a high-speed belly dip into a river on a hot day to wet its belly
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
