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Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Golden eagle (Aquila chrysaetos) adult female taking off, flying over mountain landscape, trained bird photographed during filming in Glenveagh National Park, Donegal, Ireland. August 2010
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Blackbird (Turdus merula) male in flight during snowfall, Oisterwijk, The Netherlands. January
Long eared owl (Asio otus) in flight, taking off from oak tree snag. Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Steppe eagle (Aquila nipalensis) in flight, Oman, November
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Small emperor moth (Saturnia pavonia) male below female both displaying eyespots on Gorse, Captive, UK, April
Siberian Jay (Perisoreus infaustus) alighting on a branch. Hamra, Sweden, September
Yellowhammers (Emberiza citrinella) coming in to land of a sheaf of oats in winter. Perthshire, Scotland, February
Rhinoceros hornbill (Buceros rhinoceros) male inflight, carrying mouse prey back to nest, Budo-Sungai Padi National Park, Thailand
Ghost bat (Macroderma gigas) in flight, Pine Creek, Northern Territory, Australia
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland
Ruby-throated hummingbird (Archilochus colubris) male landing on Virginia creeper (Parthenocissus quinquefolia). Hill Country, Texas, USA
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands