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RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
RF - Eyelash Pit Viper (Bothriechis schlegelii) Costa Rica
Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
RF- Masai giraffe (Giraffa camelopardalis tippelskirchi) two young calves together, Masai Mara Game Reserve Kenya
Group of African elephants (Loxodonta africana) silhouetted at sunrise, Okavango Delta, Botswana
RF- Sumatran tiger (Panthera tigris sumatrae) with cub, aged four months, captive, occurs in Sumatra, Indonesia
RF- Portrait of Siberian tiger (Panthera tigris altaica) walking in snow, captive
RF- Young male Bengal tiger, head portrait in profile (Panthera tigris tigris). Bandhavgarh National Park, India
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
RF- Great grey owl (Strix nebulosa) in flight, Finland, March
RF- Razorbill (Alca torda) resting on rocks. Great Saltee Island, County Wexford, Ireland. April
White tailed eagle (Haliaeetus albicilla) in flight with snowy background, Rausu, Japan, February
RF - Head portrait of Spotted eagle owl (Bubo africanus), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February 2016
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
RF- Eurasian Eagle owl (Bubo bubo) adult perched on rocky outcrop with the Super Full Moon, September 28th 2015, Southern Norway. August. Multiple exposure
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Buff-tailed Bumblebee (Bombus terrestris) worker feeding on Ox-eye Daisy (Leucanthemum vulgare). Peak District National Park, Derbyshire, UK. July
RF - Red Squirrel (Sciurus vulgaris) adult in summer coat leaping onto fallen log. Scotland, UK. September
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Red Squirrel (Sciurus vulgaris) adult sitting and feeding in snowfall. Cairngorms National Park, Scotland, UK, March
RF- Common frog (Rana temporaria) portrait, Cornwall, UK, January
RF- Serotine bat (Vespertilio serotinus) in flight above Hogweed. Captive, UK
RF- Red squirrel in autumn (Sciurus vulgaris). Strathspey, Scotland, UK
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Brown-lipped snail (Cepaea nemoralis) on fungus, reproduced in a bokeh affect, Sheffield, England, UK. November. Commended in the Natures Studio Category of GDT competition 2014
RF- Hyacinth macaw (Anodorhynchus hyacinthinus) in its nest hole. Pousada Aguape Lodge, Mato Grosso do Sul State, Brazil. September
RF - Siberian tiger (Panthera tigris altaica) in snow, captive
Wild horses / Mustangs, two stallions play fighting, Pryor Mountains, Montana, USA, July 2010
RF - Granular poison frog (Oophaga granulifera). Osa Peninsula, Costa Rica. Vulnerable species
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
Tail fluke of a diving Humpback whale (Megaptera novaeangliae) misty coast in background, Alaska, USA. Gulf of Alaska, Pacific ocean
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
RF- Grey seal (Halichoerus grypus) pups on Norfolk Beach, England, UK, December
RF- European hedgehog (Erinaceus europaeus) orphan, Jarfalla, Sweden. August
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
