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Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November
Huge iceberg with streaked sky under expansive open sky. Drake Passage, near the South Shetland Islands, Antarctica, December
Iceberg off the Antarctic Peninsula, Antarctica, February 2009, Taken on location for BBC Frozen Planet series
Brown hare (Lepus europaeus) running past hay barn in snow storm, Lapua, Finland, March. WINNER, Animals in their Environment category, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR competition
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
Common Swallowtail butterfly (Papilio machaon) resting on Meadow Clary (Salvia pratensis) flowers, on a white background. Nordtirol, Tirol, Austrian Alps, Austria, 1700 metres altitude, July
Yorkshire Terrier, with Netherland-cross rabbit
Young windmill-eared rabbit and matching guinea-pig
Two ginger kittens rolling playfully on their sides
Blue Bearded Collie puppy, 3 months, yawning
Two ginger kittens lying on their sides
Ginger kitten, 7 weeks, sitting between two young Lionhead-Lop rabbits
Border Collie bitch, running towards the camera
Chinchilla Persian female cat, 6 years
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Yellow Labrador Retriever puppy, 7 weeks
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
A young Spiny / Yellow / Longsnouted Seahorse (Hippocampus guttulatus), in an aquarium attached to plastic seagrass leaves. This seahorse is part of a UK seahorse breeding program
West Highland White Terrier sitting
West Highland White Terrier lying
Playful Border Collie puppies, 6 weeks
Border Collie puppy portrait
Border Collie having a good shake
Border Collie with one of her puppies, 5 weeks, yawning
Conifer trees laden with snow in winter landscape, Kuusamo, northern Finland, February 2009
Common raven (Corvus corax) perched in snow beside birch tree, Finland, March
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June
Konik Wild Horse (Equus ferus caballus) in grasslands with Purple loosestrife flowers (Lythrum salicaria) in foreground. The Netherlands, July
Domestic dog, Scottish Terrier / Aberdeen Terrier, studio portrait
Ptarmigan (Lagopus mutus) male in flight over snow covered Cairngorm Mountains, Scotland, February. (non-ex)
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
These hatchlings of an Olive ridley sea turtle (Lepidochelys olivacea) should be buried and protected under 40 cm of warm sand, but a domestic dog dug them out at Ostional beach in Costa Rica
Mountain hare (Lepus timidus) running across wind swept snow field, near Carrbridge, Cairngorms National Park, Scotland, January 2010
Kromfohrlander running / racing in field towards camera
Dalmatian, head portrait, male aged 4 years
Alaskan Malamute portrait of blue coated adult
Flock of Cattle Egrets (Bubulcus ibis) in roosting trees at dusk, Imba Ndutu, Serengeti NP, Tanzania, East Africa
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
