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Grey Wolf (Canis lupus) head portrait of male, lying in snow, Captive
Jungle Crow (Corvus macrorhynchos) perched on tree branch with rainbow, Shejila mountain, Qinghai-Tibetan Plateau, Tibet, China. July
Drystone walls near Litton, Peak District National Park, Derbyshire, UK, May
Thawing alpine lake, Vanoise National Park, Rhone-Alpes, France, June
RF- Pine marten (Martes martes) in rain. Scotland, UK. July 2015
View looking up to the summit of K2 (8, 611m), Central Karakoram National Park, Pakistan, June 2007
Whooper swan (Cygnus cygnus) in flight in snowfall, Lake Kussharo, Japan, February
White tailed eagle (Haliaeetus albicilla) in flight, Rausu, Japan, February
Arable fields in snow, near Southrepps village, North Norfolk, January
White rhino (Ceratotherium simum) in the rain, Nakuru National Park, Kenya
Snowy owl (Bubo scandiaca) female in flight, Canada. February
Detail of Cumulonimbus cloud at sunset. Nordtirol, Austrian Alps. July
Fallow deer heard (Dama dama) in heavy snow. Chatsworth Park, Derbyshire, Peak District National Park. January
Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February
Gannet (Morus bassanu) coming into land with stormy seas behind, Shetland Islands, Scotland, UK, September. Highly Commended in the Coastal and Marine category of the BWPA Awards 2015
Cliff top view of a gannet (Morus bassanus) colony. Shetland Islands, Scotland, UK, July
Osprey (Pandion haliaetus) with fish prey on feeding perch in the rain, Cairngorms National Park, Scotland, UK, July
Robin perched on boot {Erithacus rubecula} UK
Robin (Erithacus rubecula) in snow, Titchwell, Norfolk, January. Image digitally manipulated to add more snow
Robin (Erithacus rubecula) on snow covered ground, during snowfall. Derbyshire, UK. January. (non-ex)
Common magpies (Pica pica) perched on frost covered branches, Jvaskyla, Finland, January
Orcas (Orcinus orca) pod feeding on herring, wide shot showing surrounding landscape, Iceland, January 2013
Long-tailed / Crab-eating macaque (Macaca fascicularis) foraging on coastline at low tide, Bako National Park, Sarawak, Borneo, Malaysia
Snowy Owl (Bubo scandiaca) flying low, captive, February
Sossusvlei dunes, wind in rainy season, Namib-Naukluft National Park, Namib desert, Namibia, April 2008
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Sunset clouds over Paine mountains, seen from Pehoe Lake, Torres del Paine National Park, Patagonia, Chile
Brown hare (Lepus europaeus) running past hay barn in snow storm, Lapua, Finland, March. WINNER, Animals in their Environment category, 2011 WILDLIFE PHOTOGRAPHER OF THE YEAR competition
Snow falling on arable land with Oak trees in the background, Southrepps, Norfolk, UK, April
The isthmus between Greater and Little Sark, bound in snow. The heaviest snowfall on Sark in living memory. Channel Islands, UK, December 2010
Dramatic seas crashing onto a rocky shore. The Broken Group Islands from The Wild Pacific Trail, Ucluelet, West coast of Vancouver Island, Canada, September 2010
Brown bear (Ursus arctos) strolling along the beach in the rain, looking towards the camera. Lake Clark National Park, Alaska, USA, July
Dramatic light on Benijo Beach. Anaga Peninsula, Tenerife, Canary Islands, February 2011
Black browed albatross (Thalassarche melanophrys) flying in 60 knot wind at Cape Horn, Chile
Morraine Lake, in the Valley of the Ten Peaks, after recent snowfall, Banff National Park, Alberta, Canada. October 2009
The Valley of the Ten Peaks, after recent snowfall, Banff National Park, Alberta, Canada. October 2009
Storm approaching beach at Cape Trafalgar, Canos de Meca, Cadiz, Spain
Clouds fill the valley of Llobegat in Cadi Moixero Natural Park. Catalonia, Pyrenees, Spain
Moose (Alces alces) bull in snow storm with aspen trees in background, Grand Teton NP, Wyoming, USA
Lighting strikes the Andes, Altiplano region, SW Bolivia, South America
Dust storm Kalahari Gemsbok NP South Africa, summer
Green turtle (Chelonia mydas) resting in the shallows of the coast, Bijagos Islands, Guinea Bissau. Endangered species
Aerial view of Amazon Rainforest, and the Manati River, Peru, July 2015
King Penguin chicks (Aptenodytes patagonicus) huddling together in blowing snow for safety as they await the return of their parents from the sea. Gold Harbor, South Georgia, South Atlantic Islands
Mist over Coniston Water before dawn. Lake District, Cumbria, England
Small path in frosty landscape, Vosges, France, November 2013
Aurora Borealis display over mountains at night, West Icelend, January 2013
Banner cloud formation around the summit of the Matterhorn (4, 478m), Switzerland, September 2011