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Gannet (Morus bassanu) coming into land with stormy seas behind, Shetland Islands, Scotland, UK, September. Highly Commended in the Coastal and Marine category of the BWPA Awards 2015
Cliff top view of a gannet (Morus bassanus) colony. Shetland Islands, Scotland, UK, July
Osprey (Pandion haliaetus) splashing water as it emerges from loch whilst fishing at dawn, Cairngorms National Park, Scotland, July
Osprey (Pandion haliaetus) at surface of a loch after diving for a fish, Cairngorms National Park, Scotland, UK, July
Kingfisher (Alcedo atthis) flying up from water with caught fish, Worcestershire, UK, March
Golden damselfish (Amblyglyphidodon aureus) hiding amongst the branches of red whip coral (Ctenocella sp.) Misool, Raja Ampat, West Papua, Indonesia
Nudibranch (Flabellina lineata) in aquarium, taken in the field in Gulen, Norway. North East Atlantic Ocean. Collected for scientific research
Tiny (10mm) pygmy seahorse (Hippocampus bargibanti) sheltering in seafan (Muricella sp)Tiny (10mm) pygmy seahorse (Hippocampus bargibanti) sheltering in seafan (Muricella sp.) Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Portrait of Red Sea anemonefish (Amphiprion bicinctus) against open water. Nuweiba, Sinai, Egypt
Eider (Somateria mollissima) male in flight in evening light. Aberdeenshire, Scotland, UK, March
Close up detail photo of a Spotted moray (Gymnothorax isingteena), Baa Atoll, Maldives. Indian Ocean
Cleaner wrasse (Labroides dimidiatus) swims out from the gill of a Blue spotted pufferfish (Arothron caeruleopunctatus). East of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Clarks anemonefish (Amphiprion clarkii) portrait in its host Bubble-tip anemone: Entacmaea quadricolor). Horseshoe Bay, Rinca Island, Komodo National Park, Indonesia
Vibrant Red Sea reef scene, with orange female Scalefin anthias (Pseudanthias squamipinnis) swarming in front of Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents
An adult Coot (Fulica atra) running across the surface of a lake, Derbyshire, England, UK, March 2010. Did you know? It is a current theory that dinosaurs such as Archaeopteryx developed wings to
Siskin (Carduelis spinus) male reflected in garden pool. Scotland, May
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Blue shark (Prionace glauca) near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
Group of Circular spadefish (Platax orbicularis) gather at a cleaning station beneath a table coral (Acropora sp.) on a coral reef. Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
RF - Roving coralgrouper (Plectropomus pessuliferus) is cleaned by Bluestreak cleaner wrasse (Labroides dimidiatus). Ras Mohammed Marine Park; Sinai, Egypt. Red Sea
Green jacks (Caranx caballus) long exposure, La Paz, Mexico. Gulf of California, Sea of Cortez, East Pacific Ocean
Red gorgonians (Lophogorgia chilensis) growing beneath a Giant kelp (Macrocystis pyrifera) forest, with female California sheephead (Semicossyphus pulcher). Santa Barbara Island, Channel Islands
Amphipod travelling on sea gooseberry (Ctenophora) Browning Pass, Vancouver Island, British Columbia, Canada. North East Pacific Ocean
Portrait of a Nudibranch (Flabellina pellucida) on a kelp frond with star trails. Gulen, Norway. North East Atlantic Ocean
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
Grey seal (Halichoerus grypus) young female opens her mouth and blows bubbles through her nose playfully while she looks up from the seabed. Lundy Island, Devon, UK, Bristol Channel. August
RF - Scissortail sergeants fish (Abudefduf sexfasciatus) sheltering beneath hard coral (Acropora sp)RF - Scissortail sergeants fish (Abudefduf sexfasciatus) sheltering beneath hard coral (Acropora sp.) in the evening on a coral reef, Red Sea
Nudibranch (Polycera quadrilineata) on kelp fronds. Gulen, Norway. North East Atlantic Ocean
Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean
Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp)Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei)
Photo showing detail of the underside and mount of a honeycomb sea star (Pentaceraster alveolatus) with two tiny shrimps (Periclimenes soror). Bitung, North Sulawesi, Indonesia
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Redtail butterflyfish (Chaetodon collare) pair in front of a smooth Sea fan (Annella mollis). Butterflyfish regularly form stable pairs. Similan Islands, Thailand. Andaman Sea, Indian Ocean
RF - Silky shark (Carcharhinus falciformis) swimming beneath the surface of the ocean, with evening sun burst. Gardens of the Queen National Park, Cuba
RF - Galapagos sea lions (Zalophus wollebaeki) five females hauled out on a beach. Espagnola Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
Spotlight goby (Gobiosoma louisae) looking out from inside a Yellow tube sponge (Aplysina fistularis), growing on a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands, British West Indies
Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply
Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Imitator damselfish (Pomacentrus imitator) male, abstract in in front of a Black coral tree (Antipathes sp.) on a coral reef. Ra Province, Viti Levu, Fiji, Polynesia
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau
Portrait of a male River blenny (Salaria fluviatilis) River Flumendosa, Gennargentu National Park, Sardinia, Italy
Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August
