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Yesnaby sea stack in stormy light, Orkney, Scotland, October 2014
Osprey (Pandion haliaetus) fishing, Rothiemurchus, Cairngorms National Park, Scotland, UK, July
Old Mann of Storr at dawn, Isle of Skye, Inner Hebrides, Scotland, UK, April 2014
Red Cuillin in evening light from Elgol, Isle of Skye, Inner Hebrides, Scotland, UK, October 2014
Abstract of waterfall with Silver birch (Betula pendula) above, Glenfeshie, Cairngorms National Park, Scotland, UK, October 2014
Rock archway at sunset, Isle of Lewis, Outer Hebrides, Scotland, UK, September 2014
View across Sound of Taransay to North Harris hills in stormy weather, West Harris, Outer Hebrides, Scotland, UK, September 2014
Two Bottlenose dolphins (Tursiops truncatus) breaching, Moray Firth, Scotland, UK, July 2014
Bottlenose dolphin (Tursiops truncatus) breaching, Moray Firth, Scotland, UK, July
Osprey (Pandion haliaetus) fishing, about to seize prey, Rothiemurchus, Cairngorms National Park, Scotland, UK, July
Ben Mor Coigach and Loch Dhonnachaidh in winter, Assynt, Highlands, Scotland, UK, January 2014
Rock formations on Bhudhie Bheinn lit by the setting sun. Knoydart, Scotland, UK, June 2016
Rising sun illuminating Luinne Bheinn with the river Barrisdale in the forground. Knoydart, Scotland, UK, June 2016
Loch Hourn and big skies from Ladar Bheinn. Knoydart, Highlands, Scotland, UK, June 2016
Mauve stinger (Pelagia noctiluca) at the entrance of a cave, Sula Sgeir, Outer Hebrides / Western Isles, Scotland, July
Atlantic puffin (Fratercula arctica) diving, North Rona, Outer Hebrides / Western Isles, Scotland, UK, July
Great Skua (Stercorarius skua) silhouetted at sunrise, Fair Isle, Shetland, Scotland, UK, June
View over Uath Lochans before sunrise following heavy snowfall, Cairngorms National Park, Scotland, UK, November 2015
European river otter (Lutra lutra) swimming over kelp, Shetland, Scotland, UK, April
Alt Ruadh river flowing through woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, October 2013
Aerial view over Insh Marshes National Nature Reserve, Cairngorms National Park, Scotland, UK, May 2016
Waves crashing on Loch Bad a Ghaill, Assynt, Highlands of Scotland, UK, January 2016
Fionn Loch at sunset. Great Wilderness, Highlands of Scotland, UK, January 2016
Devils Point and River Dee at sunrise, in snow, Cairngorms National Park, Highlands of Scotland, UK, January 2016
Yellow stripe clingfish (Diademichthys lineatus) living in association with the Fire Urchin (Astropyga radiata) Lembeh Strait, North Sulawesi, Indonesia
Lower Falls at Artist Point, Yellowstone River. Yellowstone National Park, Wyoming, USA, June
Erect-crested penguins (Eudyptes sclateri) bathing and preening prior to moulting. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Polar bear (Ursus maritimus) sow with two juveniles walk along the ice edge during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Juvenile Banggai cardinalfish (Pterapogon kauderni) with sea anemone (Heteractis magnifica). Lembeh Strait, North Sulawesi, Indonesia
Juvenile Green turtle (Chelonia mydas) Armenime cove, South Tenerife, Canary Island, Atlantic Ocean
American flamingo (Phoenicopterus ruber) at edge of water, Borrero Bay, Santa Cruz Island, Galapagos, Ecuador
Blue-footed booby (Sula nebouxii) plunge-diving at high speed, San Cristobal Island, Galapagos, Ecuador
Fulmars (Fulmarus glacialis) on water, feeding at a fish processing plant outfall. Grundafjordur, Iceland, March
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Aerial view of Tiputini River and surrounding Rainforest. Yasuni National Park, Amazon Rainforest, Ecuador, South America
Canoes on the moored on the Nam Song River at Vang Vieng, Laos, March 2009
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
Young Baikal seal (Pusa sibirica) at breathing hole. Lake Baikal, Russia, April
The Old Man of Storr at dawn, the Isle of Skye, Scotland, UK. December 2013
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean
Silvertip sharks (Carcharhinus albimarginatus) in open water, Revillagigedo islands, Mexico. Pacific Ocean
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Amphipod (Amphipoda) living inside sea squirt (Rhopalaea sp. ) Anilao, Batangas, Luzon, PhilippinesAmphipod (Amphipoda) living inside sea squirt (Rhopalaea sp.) Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Okavango robbers (Rhabdalestes maunensis) fish swimming up the temporary Selinda Spillway, northern Botswana. This shallow river cuts across the Kalahari Desert
Jaguar (Panthera onca) drinking from the rivers edge, Mato Grosso, Pantanal, Brazil. August
Fin whale (Balaenoptera physalus) surfacing with mountain landscape, Liefdefjorden, Svalbard, Norway. July 2011