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Kermode Bear (Ursus americanus kermodei) walking near a salmon stream in the early evening hours, Great Bear Rainforest, British Columbia, Canada
Polar bear (Ursus maritimus) young male resting on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Dorsal fin of a Basking shark (Cetorhinus maximus) at the surface, Cairns of Coll, Inner Hebrides, Scotland. UK, June
Great crested grebe (Podiceps cristatus) pair of adults during part of their elaborate courtship ritual during the very last few rays of evening sunlight, Derbyshire, UK, March
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Sperm whale (Physeter macrocephalus) tail fluke above water during dive, Kaikoura, New Zealand, July, Vulnerable species
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in March. Wapusk National Park. Churchill, Manitoba, Canada
Vancouver Island black bear (Ursus americanus vancouveri) mother with cubs on a beach, Vancouver Island, British Columbia, Canada, July
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
O Shaughnessys dwarf iguana / Red-eyed dwarf iguana (Enyalioides oshaughnessyi) portrait, Mashpi, Pichincha, Ecuador, Vulnerable species
Pebas stubfoot toad / Harlequin toad (Atelopus spumarius) stretching on branch, Yasuni National Park, Orellana, Ecuador, Vulnerable Species
Yellow footed tortoise (Chelonoidis denticulata) portrait, Yasuni National Park, Orellana, Ecuador. Vulnerable species
Coastal Ecuador smoky jungle frog / Choco jungle-frog (Leptodactylus peritoaktites) sitting on stone in stream, Canande, Esmeraldas, Ecuador, Vulnerable species
Stellers Sea Eagles (Haliaeetus pelagicus) fighting on ice floe, Hokkaido, Japan. February
Stellers Eagle (Haliaeteus pelagicus) close up of head and beak, Hokkaido, Japan, February
Whale shark (Rhincodon typus) silhouette, feeding on floating fish eggs (not visible) just below the surface in calm weather. Note the bow wave ahead of the shark
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
Whitetip shark (Carcharhinus longimanus), accompanied by Pilotfish (Naucrates ductor), swims towards the camera as it cruises beneath the surface of the Red Sea, close to Little Brother Island
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
Galapagos land iguana (Conolophus subcristatus) skin and scales close-up. Isabela Island, Galapagos, Ecuador, June
Marine iguanas (Amblyrhynchus cristatus) basking on volcanic rock, Punta Espinosa, Fernandina Island, Galapagos Islands, East Pacific Ocean
Adult Giant Anteater (Myrmecophaga tridactyla) walking across savannah. Los Llanos, Colombia, South America
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Verreauxs sifaka (Propithecus verreauxi) running across a road with bus in background, Berenty Private Reserve, southern Madagascar, August 2016
Great white shark (Carcharodon carcharias) Guadalupe Island, Mexico, Pacific Ocean
Humboldt penguin (Spheniscus humboldti) group gathering at landing site. Tilgo Island, La Serena, Chile. Vulnerable species
Loggerhead turtle (Caretta caretta) hatchling running to the sea, lycian coast, Mediterranean Sea, Turkey
Polar bear (Ursus maritimus) investigating camera and batting it with paw, on pack ice off the 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska, Beaufort Sea
Olive Ridley Sea Turtle (Lepidochelys olivacea) nesting, Arribada (mass nesting event), Playa Morro Ayuta, Oaxaca state, southern Mexico. Vulnerable species
Tawny nurse shark (Nebrius ferrugineus) Vaavu Atoll, Maldives, Indian Ocean
Split level photo of an American crocodile (Crocodylus acutus) floating at the surface over a shallow seagrass meadow, close to mangroves
Rio Jatuntianhua Robber Frog (Pristimantis eriphus) male on a leaf. Ecuador, Vulnerable species
Thomass leaf monkey (Presbytis thomasi) Geunung leuser NP, Sumatra
Polar bear (Ursus maritimus) underwater view swimming in a pool, Anchorage Zoo, Alaska, USA. No release available
Silver tabby kitten with paws raised