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American crocodile (Crocodylus acutus) shows off its teeth. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Painted mantella frog (Mantella madagascariensis) Mantadia NP, Eastern Madagascar
Polar bear (Ursus maritimus) walking on Champ Island glacier above the sea, Franz Josef Land, Russian Arctic. Highly commended in Animals in their Environment Category of the Wildlife Photographer of
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Shoebill stork (Balaeniceps rex) portrait. Swamps of Mabamba, Lake Victoria, Uganda
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
Rescued and orphaned Grey-headed Flying-fox (Pteropus poliocephalus) in captivity, feeds on the nectar of a flowering native eucalyptus tree. ÔÇ¿Black Rock, Victoria, Australia
Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Polar bear cubs (Ursus maritimus) triplets age 2-3 months next to their mother. Wapusk National Park, Manitoba, Canada. February
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Giant anteater (Myrmecophaga tridactyla) Serra de Canastra National Park, Brazil
Yellow-headed Picathartes / White necked rockfowl (Picathartes gymnocephalus) Kambui Hills, Sierra Leone, Vulnerable species. Photograph taken on location for BBC Africa series, September 2010
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
Great Bustard (Otis tarda) males displaying, Salisbury Plain, Wiltshire, UK, April
Polar bear (Ursus maritimus) tagged female with wet matted fur walks over newly forming pack ice, Beaufort Sea, offshore from the 1002 area of the Arctic National Wildlife Refuge, Alaska
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
Black crowned-crane (Balearica pavonina) foraging in wetland area, Masai Mara, Kenya
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Red panda (Ailurus fulgens) captive, occurs in China
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
Southern cassowary (Casuarius casuarius), adult male with chicks keeping a wary eye on the photographer. Near Bloomfield, north Queensland, Australia, September
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
King cobra (Ophiophagus hannah) in strike pose, Malaysia