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Six-spot burnet moth (Zygaena filipendulae) resting on Red Clover (Trifolium pratense) Kingcombe Meadows DWT reserve, Dorset, UK, May
Bed of Common mussels (Mytilus edulis) growing on rocks, feeding, Megavissey, Cornwall, UK, May
Leachs spider crab (Inachus phalangium) sheltering from the current beneath tentacles of Snakelocks anemone (Anemonia viridis) Selsey, West Sussex, UK, May
Common darter dragonfly (Sympetrum striolatum) adult female at rest on Bell heather (Erica cinerea), Minsmere RSPB reserve, Suffolk, UK, July
Modern wind pump for pumping water onto Wicken Fen, Charlock (Sinapis arvensis) flwoering in the foreground, Wicken Fen, Cambridgeshire, UK, June 2011
Ringlet butterfly {Aphantopus hyperantus} resting on yellow rattle flowers, Denmark Farm, Lampeter, Wales, UK. June
RF- Large red damselfly (Pyrrhosoma nymphula) emerging from nymphal case. Cornwall, UK. April
Rear view of Black grouse (Tetrao tetrix) male displaying at lek, Cairngorms National Park, Grampian, Scotland, UK, April
Rhyne (drainage ditch) on Butleigh Moor, Somerset Levels, Somerset, England, UK, June 2011
Pussy willow catkin (Salix caprea) bursting out from bud, Somerset Levels, Somerset, England, UK, April
European brown hare (Lepus europaeus) adult grooming beside field of rapeseed, Hope Farm RSPB reserve, Cambridgeshire, UK, May
Wild boar (Sus scrofa) female in woodland undergrowth sniffling air for scent of human, Forest of Dean, Gloucestershire, UK, March, Highly commended, Animal Portraits
Red deer (Cervus elaphus) stag bellowing during autumn rut, Richmond Park, London, England, UK, October
Birch trees blowing in high winds, long exposure, Calke Abbey, The National Forest, Central England, UK, November 2010
European storm-petrel (Hydrobates pelagicus) in flight over Irish sea off Pembrokeshire coast, Wales, UK, August 2010
Dipper (Cinclus cinclus) portrait, standing on exposed stone in fast flowing river, with food for chicks in beak, Brecon Beacons National Park, Wales, UK
White Japanese Spitz dog, Sushi, 6 months old, holding a red rose
Jack-in-a-bucket - two Jack Russell Terrier puppies in a Halloween pumpkin bucket, one smooth coated, one rough coated
Black kitten in garden, Germany
Tabby kitten, Stanley, 4 months old, breaking through paper
Tabby male kitten, Fosset, 4 months old, lying with his head up
Ginger kitten, Tom, 3 months, looking over his shoulder, wearing a Sombrero hat, against white background
Tabby kittens, Stanley and Fosset, 12 weeks, walking together in unison
Colourpoint kitten in a tea cup
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Cowberry or Lingonberry (Vaccinium vitis-idaea) close-up of red berries in late summer, Scotland, UK, August
White Water Lily (Nymphaea alba) in flower, Cairngorms National Park, Scotland, UK
Common buzzard (Buteo buteo) perching on gate in falling snow, Scotland, UK, January
Eurasian otter (Lutra lutra) portrait, Scotland, UK, April
Sea stacks off Mangurstadh / Mangersta Beach, Isle of Lewis, Outer Hebrides, Scotland, UK, March 2015
Red squirrel (Sciurus vulgaris) on tree trunk, Cairngorms National Park, Scotland, UK, August
Nyiragongo volcano lava lake, Virungas National Park, Democratic Republic of Congo February 2015
Black Grouse (Tetrao tetrix) male displaying with breath condensing in cold air, Kuusamo, Finland April
Quaking aspen trees (Populus tremuloides) and conifers with dusting of snow, Dome Hill above Dawson City, Yukon Territories, Canada, September 2013
European hornet (Vespa crabro) worker in flight, Surrey, England, August
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Black eared white Boxer puppy, lying with head up and crossed paws, against white background
Chocolate Labradoodle puppy, 9 weeks, sitting and looking up, against white background
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
Temperate rainforest with Vine maple (Acer circinatum) and fern, Golden Ears provincial park, British Columbia, Canada, July
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Pika (Ochotona princeps) in scree rock pile, Sheepeaters Cliff, Yellowstone National Park, Wyoming, USA, September
Harvest mouse (Micromys minutus) on wheat stem feeding, Devon, UK, July. Captive
Pine marten (Martes martes) juvenile male sitting on branch of a dead tree outside an old black woodpecker (Dryocopus martius) nest hole. Molde, Central Norway, September
Paradise Riflebird (Ptiloris paradiseus) young males size each other up between practice displays. Australia
Lily of the valley (Convallaria majalis), Foret de Puvenelle, Lorraine Regional Natural Park, Lorraine, France, May
Male Rock ptarmigan (Lagopus mutus) vocalising whilst in flight, showing winter plumage, Utsjoki, Finland, April
