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Seven Sisters chalk cliffs, South Downs, England
Eddies in the River Sligachan with Marsco in background. Isle of Skye, Scotland, UK, October 2011
View over Glen Avon to Beinn Mheadhoin in winter, Cairngorms National Park, Scotland, December 2011
European Hare (Lepus europaeus) running. Wales, UK, July
Great spider crab (Hyas araneus) on Deadmans fingers (Alcyonium digitatum) coral, feeding in a current, Loch Carron, Ross and Cromarty, Scotland, UK, April
European Hare (Lepus europaeus) courting pair in grassland at dawn. Wales, UK, July
Water Vole (Arvicola terrestris) eating, Kent, England, UK, March
Bog myrtle / Sweet gale (Myrica gale) flowers, Caesars Camp, Fleet, Hampshire, England, UK, March
Flowering Heather and Bracken on lowland heath, with path in the background, Caesars Camp, Fleet, Hampshire, England, UK, August
Recently burnt heathland, showing new Bracken growth (Pteridium aquilinum), Caesars Camp, Fleet, Hampshire, England, UK, May
Close-up of flowering Bell Heather (Erica cinerea), Caesars Camp, Fleet, Hampshire, England, UK, August
Field of ripe Oats with Combine harvester in the distance, Haregill Lodge Farm, Ellingstring, North Yorkshire, England, UK, August
Wood Anemone (Anemone nemorosa) in flower. Powerstock Common Nature Reserve, Dorset, England, UK, April
Male Blackbird (Turdus merula) perched on old barn roof, Inverness-shire, Scotland, UK, November
Capercaillie (Tetrao urogallus) male displaying. Cairngorms National Park, Scotland, UK, September
Rowan Tree (Sorbus aucuparia) covered in berries seen through autumn leaves. Dartmoor National Park, Devon, England, UK, October
Peregrine Falcon (Falco peregrinus) in flight over the River Tay. Kinnoull Hill Woodland Park, Perthshire, Scotland, November
Mountain Hare (Lepus timidus) in alert pose. Cairngorms National Park, Scotland, July
Whooper Swans (Cygnus cygnus), ducks and wigeon feeding in the estuary, Caerlaverock WWT, Scotland, Solway, UK, January
Worker honey bee (Apis mellifera) feeding on nectar of Borage (Borago officinalis) flower in conservation margin at RSPBs Hope Farm, Cambridgeshire, UK. May
Short-eared owl (Asio flammeus) perched on a fence post, Worlaby Carr, Lincolnshire, England, UK, December
Young Red fox (Vulpes vulpes) walking over ice of frozen pond in garden, Bristol, UK, February
Portrait of a young Grey seal (Halichoerus grypus) Farne Islands, Northumberland, UK, North Sea, July
Red squirrel (Sciurus vulgaris) on branch in morning sun, Brownsea Island, Dorset, UK, February
Mute Swan (Cygnus olor) standing on ice at sunrise. Glasgow, Scotland, December
Northern wheatear (Oenanthe oenanthe) adult male in spring plumage feeding on dung flies at farm midden heap, Hertfordshire, UK, April
Common dolphin (Delphinus delphis) bow-riding, near South Uist, Outer Hebrides, Scotland, UK, June
Mixed flock of Northern gannets (Morus bassanus) and gulls (Larus) feeding in the wake of the pelagic trawler Charisma in evening light, Shetland Islands, Scotland, UK, October 2011
Harvestman (Opiliones) on flowering heather, Arne RSPB reserve, Dorset, England, UK, July
Common starling (Sturnus vulgaris) adult with insect food arriving at nestbox to feed chicks, Hope Farm RSPB Reserve, Cambridgeshire, England, UK, May
Seven spot ladybird (Coccinella septempunctata) on Black knapweed (Centaurea nigra) flower at RSPBs Hope Farm Reserve, Cambridgeshire, England, UK, July
Two young Coots (Fulica atra) on a still lake at dawn, Derbyshire, England, UK, June 2010
Portrait of a male Roe deer (Capreolus capreolus) in a meadow, Cairngorms NP, Scotland, UK, August 2010
Greylag goose (Anser anser) pair on edge of loch, near nesting site, Scotland, UK, May 2010
Long exposure of sea, with rocks in foreground, Bagh Dhail Mor, Isle of Lewis, Outer Hebrides, Scotland, UK, October 2011
Rock ptarmigan (Lagopus mutus) close-up portrait in winter plumage, Cairngorms NP, Scotland, UK, February
Adult male Red Grouse (Lagopus lagopus scoticus) in snow, Cairngorms NP, Scotland, UK, February
Red deer (Cervus elaphus) stag in pine woodland in winter, Cairngorms National Park, Scotland, UK, February
Great crested grebe (Podiceps cristatus) pair of adults during part of their elaborate courtship ritual during the very last few rays of evening sunlight, Derbyshire, UK, March
Dartford warbler (Sylvia undata) male perched, singing, Wales, UK, May
Greater sundew (Drosera anglica) close-up, Flow Country, Sutherland, Highlands, Scotland, UK, July
Bog Asphodel (Narthecium ossifragum) and Cross-leaved Heath (Erica tetralix) in flower, New Forest, Hampshire, England, UK, July
Bell heather (Erica cinerea) in bloom on lowland heathland, Caesars Camp, Aldershot, Hampshire, England, UK, August 2011
Great skua (Stercorarius skua) on water, Shetland Islands, Scotland, UK, April
Six-spot burnet moth (Zygaena filipendulae) resting on Red Clover (Trifolium pratense) Kingcombe Meadows DWT reserve, Dorset, UK, May
Bed of Common mussels (Mytilus edulis) growing on rocks, feeding, Megavissey, Cornwall, UK, May
Leachs spider crab (Inachus phalangium) sheltering from the current beneath tentacles of Snakelocks anemone (Anemonia viridis) Selsey, West Sussex, UK, May
Common darter dragonfly (Sympetrum striolatum) adult female at rest on Bell heather (Erica cinerea), Minsmere RSPB reserve, Suffolk, UK, July