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Portrait of a Rough Collie puppy, 14 weeks
Bearcoat Shar Pei mother, with her Blue Bearcoat puppy, 13 weeks
Portrait of a young Badger (Meles meles)
Merle Border Collie puppy, 6 weeks, lying with head up
Black kitten, 7 weeks, rolling on its back
Fawn Pug puppy, 8 weeks, wearing a Father Christmas hat
Cream Dapple Miniature Long-haired Dachshund puppy with British shorthair red tabby Kitten
Portrait of a silver tabby-and-white cat walking
Yellow Labrador retriever puppy lying with paws crossed, 8 weeks
Sable Border Collie puppy, 9 weeks, looking over his shoulder
Chocolate registered Border Collie dog, 9 months
White-faced black-and-white Border Collie puppy
Crabeater Seal (Lobodon carcinophaga) hauled out on ice. Pleneau Island, Antarctica, January
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Siberian Jay (Perisoreus infaustus) alighting on a branch. Hamra, Sweden, September
Two Common ravens (Corvus corax) in flight through snow, Finland, April
Sable Border Collie puppy, 9 weeks, sitting
Wild Boar (Sus scrofa) males fighting in snow. The Netherlands, January
Whooper Swan (Cygnus cygnus) with its beak in its feathers. The Netherlands, January
Osprey (Pandion haliaetus) flying through a rainbow. Aviemore, Scotland, July
Yellowhammers (Emberiza citrinella) coming in to land of a sheaf of oats in winter. Perthshire, Scotland, February
Domestic dog, Mongrel, studio portrait
Wolverine (Gulo gulo) climbing birch tree, Kamchatka, Far East Russia, April
A silhouette of a California sealion pup (Zalophus californianus) next to cliffs. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
Moose (Alces alces) head portrait of bull standing in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September
Hungarian Vizsla portrait amongst autumn foliage, Connecticut, USA
Golden retriever splashing through water, USA
Domestic sheep, lambs playing in a field, Norfolk, UK, March
Yellow labrador, five puppies in a row
Domestic goat (Capra hircus) pygmy goat bleeting, portrait, UK
Border collie, head portrait, Norfolk, UK, May
Aylesbury ducks following in a line on village green, Weedon, Buckinghamshire, UK, October
Mute Swan (Cygnus olor) with its beak in water. Fife, Scotland, November
Small herd of Asiatic Wild Ass (Equus hemionus) running, Gobi National Park, Mongolia
Australian Shepherd, red-tri coated, portrait sitting in snow, with picket fence behind
Miniature Schnauzer, black-silver coated, standing in autumn foliage
British Longhair Cats, tomcats, black and red coated (Highlander, Lowlander, Britannica) sitting on large log, with autumn foliage
Boston Terrier, head portrait of male
Four British Shorthair kittens, two silver-shaded and two golden-mackerel-tabby, sitting on stand
Lemon shark (Negaprion brevirostris) under sun rays. Little Bahama Bank, Bahamas. West Atlantic Ocean
Oceanic whitetip shark (Carcharhinus longimanus) with Pilot fish (Naucrates ductor). Tropical West Atlantic Ocean, off Cat Island, Bahamas
Pygmy x Golden Guernsey female goat kid, standing on hind legs
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Polar bear (Ursus maritimus) female and her two cubs on large iceberg off Baffin Island. Nunavut, Canada, August 2010
A large female Tiger Shark (Galeocerdo cuvier) swimming in open water. Grand Bahama, Bahamas. Tropical West Atlantic Ocean. March
Young Florida manatee (Trichechus manatus latirostrus) portrait, Three Sisters Spring, Crystal River, Florida, USA. February 2010
Southern stingray (Hypanus americanus) swimming over sand ripples on sandbar, Stingray City, Grand Cayman. British West Indies
Group portrait of five dogs sitting, from left to rt: two mongrels, Rough Collie, Bobtail (Old English Sheepdog) and longhaired Chihuahua