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Arctic foxes (Vulpes lagopus) juveniles playing, biting tail, Wrangel Island, Far Eastern Russia, August
Arctic fox ( Vulpes lagopus) in snow carrying snowgoose egg, Wrangel Island, Far East Russia, May
Musk ox (Ovibos moschatus) two in profile, Wrangel Island, Far Eastern Russia, September
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
Musk ox (Ovibos moschatus) at sunset, Wrangel Island, Far Eastern Russia
Polar bear (Ursus maritimus) with cub walking across ice field, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) mother with cubs walking through snow, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) group feeding on carcass on beach, Wrangel Island, Far Eastern Russia, September
Moonrise at twilight, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Waterfall during monsoon, Eravikulam National Park, Western Ghats UNESCO World Heritage Site, India, June 2012
Clouds over mountains of Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fossilized remains of Lepidendron, the tallest of the coal age trees, at the Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Iceberg in the sea outside the Ilulissat Icefjord UNESCO World Heritage Site, Greenland, August 2014. Photographed for the Freshwater Project
Aerial view of meltwater lake on the ice cap north-east of Sermeq Kujalleq Glacier, Ilulissat Icefjord UNESCO World Heritage Site, Sermersuaq / Greenland ice sheet, Greenland, August 2014
Aerial view of meltwater channels in ice cap north-east of Sermeq Kujalleq Glacier, Sermersuaq / Greenland ice sheet, Greenland, August 2014. Photographed for The Freshwater Project
Rift between the North American and Eurasian tectonic plates at Thingvellir, Thingvellir National Park, UNESCO World Heritage Site, Iceland, September. Photographed for the Freshwater Project
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
Northern Rockhopper Penguin (Eudyptes moseleyi) on nest, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Emperor shrimp (Periclemenes imperator) on sea cucumber (Thelenota ananas) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Longnose hawkfish (Oxycirrhites typus) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Travertine terraces, landscape in Huanglong Scenic and Historic Interest Area UNESCO World Heritage Sites, Sichuan, China. August. Photographed for The Freshwater Project
Jian Zhu Pu Bu (Arrow Bamboo Lake Waterfall), Jiuzhaigou National Park, Jiuzhaigou Valley Scenic and Historic Interest Area UNESCO World Heritage Site, Sichuan, China. May 2013
Travertine terraces, Huanglong Scenic and Historic Interest Area UNESCO World Heritage Sites, Sichuan, China. August. Photographed for The Freshwater Project
Fungus gnat (Mycetophilidae) larvae, with sticky hanging threads, waiting to ambush small flying insects. Deer caves, Gunung Mulu National Park, Borneo, Sarawak, Malaysia
Aerial view of melted ice around geothermal spring, Putoransky State Nature Reserve, Putorana Plateau UNESCO World Heritage Site, Siberia, Russia. June 2015
Milkweed (Asclepias mellodora), Intervales State Park, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Misty landscape in Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia. May 2014
Frozen waterfall and basalt cliffs, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Rainbow and waterfall, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Northern lights over Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Aerial view of Siberian tundra reindeer (Rangifer tarandus sibiricus) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Sunset over Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, October
Waterfall in Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, July 2014
Sable (Martes zibellina) Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Dingo (Canis lupus dingo) with Silver Gull (Larus navaehollandiae) on beach, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Lake Wabby among sand dunes and forest. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Sunrise at Seventy Five Mile Beach, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November. Model released
Sooty Tern (Sterna fuscata) in flight, Blinky Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia
Forest with mist on the slopes of Mount Lidgbird. Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia
Sooty Tern (Sterna fuscata) in flight, Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Soldiers Cap and Admiralty Islands from Malabar Hill, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
RF- Solar halo over the Laitaure delta, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September 2008
RF- Reindeer (Rangifer tarandus) portrait, semi-domesticated, small scale herding by local Sami, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September 2008
Arctic foxes (Vulpes lagopus) juveniles playing, in summer coat, Wrangel Island, Far Eastern Russia, August
Musk ox (Ovibos moschatus) portrait, Wrangel Island, Far Eastern Russia, March
