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Zapato Islet and Toro Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Mealy Parrots (Amazona farinosa) feeding at the wall of a clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Tree Fern (Cyathea sp.) detail, El Triunfo Biosphere Reserve, Chiapas, southern Mexico, May
Landscape of coniferous trees in dusk mist, Finca Nueva Linda, El Triunfo Biosphere Reserve, Chiapas, southern Mexico, April
Crested guan (Penelope purpurascens), Finca Arroyo Negro, El Triunfo Biosphere Reserve, Chiapas, southern Mexico, April
Mahogany treefrog (Tlalocohyla loquax) clasping onto leaf stem, view from below. Las Guacamayas Biological Station, Laguna del Tigre National Park, El Peten, Guatemala
Coastal landscape with empty beach Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, July
Wood Stork (Mycteria americana) feeding, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, July
Sabinal Frog (Leptodactylus melanonotus), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, July
Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site
Hyacinth macaw (Anodorhynchus hyacinthinus) pair perched on branch, silhouetted under crescent moon. Mato Grosso Forest, Pantanal, Mato Grosso, Brazil
Marsh deer (Blastocerus dichotomus) walking across track in forest. Pantanal, Mato Grosso, Brazil
Tiger (Panthera tigris tigris) female with cub, Kabini, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Mustached Bat (Pteronotus sp. ) flying at dusk, Calakmul Biosphere Reserve, Yucatan PeninsulaMustached Bat (Pteronotus sp.) flying at dusk, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
American crocodile (Crocodylus acutus) with mouth open over seagrass bed, Chinchorro Banks Biosphere Reserve, Quintana Roo, Mexico
Snow leopard (Panthera uncia) old male walking on a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains
Himalayan Griffon (Gyps himalayensis) in flight in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Snow leopard (Panthera uncia) old male at dusk looking from a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Tufted grey langur (Semnopithecus priam) baby in tree, Nilgiri Biosphere Reserve, Karnataka, India
Amazonian Horned Frog (Ceratophrys cornuta) camouflaged amongst leaf litter on lowland rainforest floor, waiting to ambush passing prey. Manu Biosphere Reserve, Amazonia, Peru. November
Katydid / Bush Cricket (Tettigoniidae) camouflaged amongst cloud forest understory vegetation. 1600 metres altitude, Manu Biosphere Reserve, Peru. November
Green-banded urania moth (Urania leilus) moths drinking salts from mineral-rich river clay, a behaviour know as puddling. On the banks of the Manu River, Manu Biosphere Reserve, Amazonia, Peru
Nictitating membrane covering eye of Map Treefrog (Hypsiboas geographicus). The membrane affords camouflage and protection from dryness whilst allowing the frog to see
River running through cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Crab Spider (Thomisidae) waiting in an orchid to ambush its prey. Cloud forest, Manu Biosphere Reserve, Peru. November
Grey whale (Eschrichtius robustus) calf, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Bell glass frog (Cochranella nola) from below, photographed on a pane of glass in Manu Biosphere Reserve, Peru
Saltpan, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, February
Snowy Egret (Egretta thula), Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Western Grebe (Aechmophorus occidentalis) swimming, Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Blurred motion image of Caribbean flamingo (Phoenicopterus ruber) juveniles, captured to be tagged and released to the wild, captive, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Black-necked Stilt (Himantopus mexicanus) walking, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
Great green bush-cricket (Tettigonia viridissima) and Common poppy (Papaver rhoeas) Sierra de Grazalema Natural Park, southern Spain, April
Dew on spiderweb with Common poppy (Papaver rhoeas) in the background, Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Dew on spiderweb with Grey-leaved cistus (Cistus albidus) Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Frost covered leaves in winter, Sierra de Grazalema Natural Park, southern Spain, February
Orchid (Orchis langei) in sunset light, Sierra de Grazalema Natural Park, southern Spain, May
Woodlark (Lullula arborea) in flowers, Sierra de Grazalema Natural Park, southern Spain, April
Common redstart (Phoenicurus phoenicurus), Sierra de Grazalema Natural Park, southern Spain, September
Great grey shrike (Lanius meridionalis) perched on a branch of Montpellier maple (Acer monspessulanum). Sierra de Grazalema Natural Park. southern Spain. December
Spanish fir tree (Abies pinsapo) covered in snow, Sierra de Grazalema Natural Park, southern Spain, November
Air bubbles in ice, Sierra de Grazalema Natural Park, southern Spain, December
Bacteria (Lepthotryx discophora) causing iridescent patterns and White willow tree leaf (Salix alba) in river, Sierra de Grazalema Natural Park, southern Spain, November