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Three-striped poison frog (Ameerega trivittata) amongst leaf litter on lowland rainforest floor. Manu Biosphere Reserve, Amazonia, Peru. November
Bush cricket or katydid (Tettigoniidae) with spines, camouflaged amongst moss. Forest interior of mid-altitude montane rainforest, Manu Biosphere Reserve, Peru
Hoatzin (Opisthocomus hoazin), pair on floating log in Cocha Salvador ox-bow lake. Manu Biosphere Reserve, Peru
Aerial view of lowland Amazonia rainforest in morning mist. Manu Biosphere Reserve, Amazonia, Peru. November 2017
Tourist touching Grey whale (Eschrichtius robustus) calf, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, February
Caribbean Flamingo (Phoenicopterus ruber) juvenile, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
Bacteria (Leptothryx discophora) causing iridescent sheen on water surface with White willow leaf (Salix alba). Sierra de Grazalema Natural Park, southern Spain, November
Blue lupin (Lupinus angustifolius) Sierra de Grazalema Natural Park, southern Spain, May
Poplar trees (Populus sp) in autumn, Sierra de Grazalema Natural Park, southern Spain, October
Hawthorn berries (Crataegus monogyna) in Poplar woodland (Populus) Sierra de Grazalema Natural Park, southern Spain. October
Frozen Elmleaf blackberry (Rubus ulmifolius) in a small pond, Sierra de Grazalema Natural Park, southern Spain, January
Leaf frozen in ice, Sierra de Grazalema Natural Park, southern Spain, February. Highly Commended in the Plants and Fungi Category of the Asferico 2016 Awards
Fog in Grazalema Mountains, Sierra de Grazalema Natural Park, southern Spain, November 2014
Firecrest (Regulus ignicapilla) perched on lichen covered branch, Sierra de Grazalema Natural Park, southern Spain, May
European bee eaters (Merops apiaster) mating, Sierra de Grazalema Natural Park, El Bosque, southern Spain, May
Giant peacock moth (Saturnia pyri) Sierra de Grazalema Natural Park, Ubrique, southern Spain, April
Golden pheasant (Chrysolophus pictus) male jumping, Yangxian Biosphere Reserve, Shaanxi, China
Red-billed blue magpie (Urocissa erythroryncha) perched on a branch, Yangxian Biosphere Reserve, Shaanxi, China, April
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Underwater view of African tigerfish (Hydrocynus vittatus), predatory fish in the Okavango river with Water lily (Nymphaea nouchali var. caerulea) and other aquatic plants
Aerial view of the meandering Saint George branch of the Danube river, Danube Delta Biosphere Reserve UNESCO World Heritage Site, Romania, May 2014. Photographed for The Freshwater Project
Golden pheasant 1+Chrysolophus pictus+2 male jumping, Yangxian Biosphere Reserve, Shaanxi, China
Red-billed blue magpie (Urocissa erythroryncha) flying, Yangxian Biosphere Reserve, Shaanxi, China
White forest crab (Cardisoma armatum) portrait, seen through claw, Island of Principe UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
Waterfall in tropical rainforest at Lacan-tun, Montes Azules Biosphere Reserve, Chiapas, Mexico
Garibaldi fish (Hypsypops rubicundus) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphere Reserve, Baja California, Mexico, May
Californian sea lion (Zalophus californianus) in Giant kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphere Reserve, Baja California, Mexico, May
Tourist boat passing the arch of Cabo San Lucas, a distinctive granitic rock formation at the southern tip of Cabo San Lucas, Baja California, Mexico, Pacific Ocean. October, 2018
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp. ) hunting a large school of Sardines (Sardina sp)Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp.) hunting a large school of Sardines (Sardina sp.), Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Group of California sea lions (Zalophus californianus) playing near the sea surface, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Two California sea lions (Zalophus californianus) playing, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
California sea lion (Zalophus californianus) close up with group playing in shallow water behind, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex dentex) hunting a huge school of Sardines (Sardina pilchardus) in shallow water, Espiritu Santo archipelago, Sea of Cortez, Mexico
Brown booby (Sula leucogaster), Los Islotes, Espiritu Santo Archipelago National Park, Sea of Cortez (Gulf of California), Mexico, September
Californian sea lion (Zalophus californianus), Los Islotes, Espiritu Santo Archipelago National Park, Sea of Cortez (Gulf of California), Mexico, September
Cliff wall and wave, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Bluehead Wrasse (Thalassoma bifasciatum) juvenile cleaning Smooth Trunkfish (Lactophrys triqueter), Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
Guadalupe fur seal (Arctocephalus townsendi) pup, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
View to the northern part of the island, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Giant coreopsis (coreopsis gigantea) flower, Zapato Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Giant coreopsis (coreopsis gigantea) inside the crater, Zapato Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Guadalupe cistanthe (Cistanthe guadalupensis) succulent, Negro Islet with main island in the back, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, September
Northern elephant seal (Mirounga angustirostris) juvenile swimming, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, September
Desert hyacinth (Triteleia guadalupensis) flower, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Zapato Islet and Toro Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Mealy Parrots (Amazona farinosa) feeding at the wall of a clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November