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Smooth-fronted caiman / Crowned dwarf caiman (Paleosuchus trigonatus) head above water, Cuyabeno, Sucumbios, Ecuador
Female Sea lion (Zalophus wollebaeki) with pup suckling in shallow water, Floreana Island, Galapagos
Werners / Tumbesian bush anole (Polychrus femoralis) on plant, Macara, Loja, Ecuador
Black caiman (Melanosuchus niger) at water surface with horse flies on its head, Yasuni National Park, Orellana, Ecuador
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Fer-de-lance (Bothrops asper) camouflaged on the rainforest floor. Corcovado National Park, Osa Peninsula, Costa Rica
Fleischmanns glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on the underside of a leaf overhanging a rainforest stream
Jumping spider (Salticidae) hunting among vegetation, San Jose, Costa Rica
Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio
Red-eyed leaf frog (Agalychnis callidryas) male, Central Caribbean foothills, Costa Rica
Granular poison frog (Oophaga granulifera) Osa Peninsula, Costa Rica. Vulnerable IUCN Red List species
Lemur leaf frog (Agalychnis lemur) at night, Central Caribbean foothills, Costa Rica. IUCN Red List critically endangered species
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Slender sapsucking slug (Thuridilla gracilis) on surface of Royal seasquirt (Polycarpa aurata) Aljui Bay, Waigeo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Harlequin shrimp (Hymenocera elegans) on blue sea star. This species of shrimps feeds on starfish, turning them upsidedown and keeping them as living larder. Bitung, North Sulawesi, Indonesia
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Hawksbill turtle (Eretmochelys imbricata) feeding on soft coral (Litophyton arboreum) Nuweiba, Sinai, Egypt. Gulf of Aqaba, Red Sea
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
Portrait of Paddle-flap scorpionfish (Rhinopias eschmeyeri) on rubble slope. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Roughhead blenny (Acanthemblemaria aspera) male portrait. This species is very small, just 20mm long and 1-2mm deep, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Secretary blenny (Acanthemblemaria maria) making a home in a deserved worm tube of a great star coral (Montastraea cavernosa), East End, Grand Cayman, Cayman Islands, British West Indies
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A Mountainous star coral (Montastraea faveolata) spawning at night, with gamete bundle being released from a coral polyp, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Whale shark (Rhincodon typus) silhouette, feeding on floating fish eggs (not visible) just below the surface in calm weather. Note the bow wave ahead of the shark
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
Whitetip shark (Carcharhinus longimanus), accompanied by Pilotfish (Naucrates ductor), swims towards the camera as it cruises beneath the surface of the Red Sea, close to Little Brother Island
Aggregation of tiny isopods (Santia sp. ) living on the surface of a Blue sponge (Haliclona sp)Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Diver exploring a freshwater cenote (or limestone sinkhole) swimming through beams of light filtered through the forest above. Garden Of Eden Cenote, Puerto Aventuras, Quintana Roo, Yucatan, Mexico
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines