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Female scalefin anthias (Pseudanthias squamipinnis) swimming in front of a school of glassfish (Apogonidae), whose shiny bodies are rendered into out of focus bokeh patterns. Laamu Atoll, Maldives
School of powderblue surgeonfish (Acanthurus leucosternon) feeding, Laamu Atoll, Maldives. Indian Ocean
Spotted Porcupine fish (Diodon hystrix) inflated, displaying defensive behaviour, North Pacific Ocean, Hawaii
Cairns birdwing butterfly (Ornithoptera euphorion) male resting on fern, large butterfly endemic to wet tropics in far north Queensland, Cairns, Queensland, Australia
Dusky leaf-nosed bat (Hipposideros ater) flying out of daytime cave roost in a sandstone escarpment, Kimberley, Western Australia
Scalloped ribbonfish (Zu cristatus) juvenile, Balayan Bay, Batangas, the Philippines
RF - Split level photo of mangrove scenery, with hard corals (including Goniopora sp. Porites sp.) growing below Red mangrove tree (Rhizophora sp.). Raja Ampat, West Papua, Indonesia
The Chocolate Hills are a geological formation in Bohol province in the Philippines. More than a thousand of these hills occur in a 50 square kilometre area
Group of Bigfin reef squids (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Bigfin reef squid (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
Queen conch (Aliger gigas) in a seagrass (Thalassia testudinum) meadow, Bahamas
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
Loggerhead turtle (Caretta caretta) accompanied by pilotfish (Naucrates ductor) swimming near the surface. Santa Maria Island, Azores, Portugal, Atlantic Ocean. Vulnerable
Atlantic spotted dolphins, (Stenella frontalis), Formigas Islet dive site, Santa Maria Island, Azores, Portugal, Atlantic Ocean
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Banded sea krait (Laticauda colubrina) hunting in seagrass bed. Manado, North Sulawesi, Indonesia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
RF - Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and Sea plumes (Pseudopterogorgia sp)
Green sea turtle (Chelonia mydas) near surface in shallow water. The Bahamas
Caribbean reef squid (Sepioteuthis sepioidea) eating fish at night, portrait. The Bahamas
School of Oriental sweetlips (Plectorhinchus vittatus) gather tightly together as they rest during the day, with a Clown triggerfish (Balistoides conspicillum)
A portrait of a clown triggerfish (Balistoides conspicillum) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Misool Marine Protected Area. Ceram Sea. Tropical West Pacific Ocean
Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea
A male pajama cardinalfish (Sphaeramia nematoptera) incubating eggs in his mouth (not clearly visible). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
British World War II BSA M20 motorbike inside Hold 2 of wreck of HMS Thistlegorm, home to pair of Red Sea soldierfish (Myripristis murdjan) and nudibranch (Nembrotha megalocera)
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos
Tail of Blue whale (Balaenoptera musculus brevicauda). This may be the pygmy sub-species of blue whale, Balaenoptera musculus. Mirissa, Sri Lanka, Indian Ocean
A reef scene with an orange elephant ear sponge (Ianthella basta) and red sea whip (Ellisella sp) with schooling silversides fish above. Pelee Island, Misool, Raja Ampat, West Papua, Indonesia
Split level photo of mangrove scenery, with hard corals ( including Goniopora sp.; Heliopora sp; Porites sp.) growing below Red mangrove tree: (Rhizophora mangle)
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Black grouper (Mycteroperca bonaci) yawning on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Spinetail devil ray (Mobula mobular), four with rays of light and sparkle of sun on water, aerial view. Sea of Cortez, Baja California, Mexico. March
RF - Open brain coral (Trachyphyllia geoffroyi) lives in symbiosis with zooxanthellae which are responsible for nutrition and the vibrant colours. Lembeh Strait, North Sulaweisi, Indonesia
Coral polyps, Honduras
White-cheeked spider monkey (Ateles marginatus), Amazon, Brazil. June
Golden cownose rays (Rhinoptera steindachneri) swimming near mangrove roots, Elizabeth Bay, Isabela Island, Galapagos
Spotted eagle rays (Aetobatus narinari) shoal among mangrove roots, Elizabeth Bay, Isabela Island, Galapagos
Green turtle (Chelonia mydas) swimming near coast with mangrove roots, , Posa de los Patillos, Fernadina Island, Galapagos
Gray snapper (Lutjanus griseus) shoal swimming over Turtle grass (Thalassia testudinum), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
American crocodile (Crocodylus acutus), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
A Bottlenose dolphin (Tursiops truncatus), heavily scarred, possibly from shark attacks, on sand bank off Eleuthera, Bahamas
A Giant clam (Tridacna gigas) surrounded by Lettuce coral (Turbinaria reniformis), Kosrae, Micronesia
