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North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October
American Buffalo or Bison (Bison bison) group with calves, Yellowstone National Park, Wyoming, USA, May
Pair of adult Trumpeter Swans (Cygnus buccinator) in flight. Skagit County, Washington. January
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Western sandpipers (Calidris mauri) roosting on barrier island tidal flats. Terrebonne Parish, Louisiana, USA. October
Spotted reed frog (Hyperolius pucticulatus) tadpoles, captive, endemic to Tanzania
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
Grey squirrel (Sciurus carolinensis) adult and baby, against white background
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
Hyacinth Macaw (Anodorhynchus hyacinthinus) small group in flight, Brazil
Japanese Macaques (Macaca fuscata) enjoy time soaking and grooming in the hot spring in Jigokudani, Japan
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Tabby kittens, Stanley and Fosset, 12 weeks old, walking together
Rothschilds giraffe (Giraffa camelopardalis rothschildi), Murchison Falls National Park, Uganda, East Africa. Murchison Falls National Park hosts the only remaining naturally occurring population of
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June
The Lovers Baobabs (Adansonia grandidieri). Near Morondava, Madagascar" The Lovers" Baobabs (Adansonia grandidieri). Near Morondava, Madagascar, Africa, September 2010
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December
Mute swan (Cygnus olor) pair on misty lake, Amsterdamse Waterleidingduinen Nature Reserve, near Zandvoort, The Netherlands, February
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
RF- Pod of Atlantic spotted dolphins (Stenella frontalis) over shallow sand bank accompanied by larger Bottlenosed dolphin (Tursiops truncatus). Sandy Ridge, Little Bahama Bank, Bahamas
Group of Whitetip reef sharks (Triaenodon obesus) resting on a ledge, Revillagigedo Islands, Mexico, Pacific Ocean
Wild Indian Ocean bottlenose dolphin (Tursiops adunctus) swimming with a golden retriever dog (Canis lupus familiaris). Gubal Island, Egypt, Red Sea, Indian Ocean
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Yellowfronted canaries (Crithagra mozambica) at waterhole, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. June
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Western Grebe (Aechmophorus occidentalis) swimming, Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Salmon (Salmonidae) fry with egg case still attached, Lake Kuril, Kamchatka, Far East Russia, January
