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Vancouver Island black bear (Ursus americanus vancouveri) mother with cubs on a beach, Vancouver Island, British Columbia, Canada, July
American Red fox (Vulpes vulpes fulva) baby leaping on its disinterested mother. Grand Teton National Park, Wyoming, USA, May
Northern raccoon (Procyon lotor), group standing on branch, captive
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
Savanna landscape wirh Grants zebras (Equus quagga boehmi) and acacia trees, Masai Mara National Reserve, Kenya, Africa
Liver Flatcoated Retriever puppy, 6 weeks, with two baby Guinea pigs
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Three Llamas (Lama glama) resting in the Machu Picchu ruins. Urubamba, Peru. August, 2016
American bald eagle (Haliaeetus leucocephalus) three perched, Homer, Alaska, Jan
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
Sunflowers (Helianthus annuus) in bloom, Norfolk, England, UK, August
Imbabura Tree Frogs (Boana picturatus) perched on a branch. Bilsa, Esmeraldas, Ecuador
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Aphelandra (Acanthus sp) flowering. Oahu, Hawaii, February
Forester kangaroo (Macropus giganteus tasmaniensis) family group, male, female and large joey, Tasmania, Australia, February. Subspecies of Eastern grey kangaroo
Black Labrador and two puppies
Star trails above Baobab trees (Adansonia za) at night, Zombitse National Park, Madagascar, June 2010
Monte Perdido massif and Las Tres Sorores (The Three Sisters) Ordesa National Park, Aragon, Huesca, Spain
Midwater pteropod molluscs / Sea butterflies (Clione limacina) Mid-Atlantic Ridge, North Atlantic Ocean
Black-tailed Prairie Dogs (Cynomys ludovicianus), two pups (right) seem to be kissing their parent. The behaviour is a gesture of recognition
Two newly fledged burrowing owl chicks (Athene cunicularia) one being groomed by its mother (far left) Pantanal, Brazil. WINNER
Mute swan (Cygnus olor) cygnets on water, Lac de Saint Point, Franche Comte, France, May
Plains zebra (Equus burchelli) three on dried mud plains, Etosha National Park, Namibia
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
African wild dog (Lycaon pictus) alpha female, interacting with two of her pups, aged 4 weeks, Okavango Delta, Northern Botswana, Africa
Three King penguins (Aptenodytes patagonicus) reflections in water, Salisbury Plain, South Georgia, Southern Ocean
Common blue butterflies (Polyommatus icarus) roosting in morning light, Devon, UK. July
Puma (Puma concolor) female with two cubs, aged six months, sitting in fresh snow, Torres del Paine National Park, Patagonia, Chile
Three Spotted hyenas (Crocuta crocuta) fighting over meat from a nearby carcass with flock of White-backed vultures (Gyps africanus) watching in background, Mana Pools National Park, Zimbabwe
Adult Guanacos (Lama guanicoe) grazing in deep snow near Lago Pehoe, Torres del Paine National Park, Chile, July
Giraffe (Giraffa camelopardalis angolensis) calf, aged 6 weeks, with two Yellow-billed oxpeckers (Buphagus africanus) perched on its head feeding on parasites, Okavango Delta, Botswana, Africa
American flamingos (Phoenicopterus ruber) wading through salt pond, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Three male Ostriches (Struthio camelus) running and flapping wings in aggressive display, Kgalagadi Transfrontier Park, South Africa
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
Female polar bear (Ursus maritimus) with two cubs walking in a line across snow, Churchill, Canada. November
Bison (Bison bison) being circled by wolves (Canis lupus) hunting in snow, Yellowstone National Park, USA. January
Three Exmoor ponies (Equus ferus caballus), semi-feral native breed, in Exmoor National Park, Somerset / Devon, England. November
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
Blue pincushion (Brunonia australis). Tasmania, Australia. December
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Three Grey partridge (Perdix perdix) walking in snow. Near Nijmegen, the Netherlands. February
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Female Galapagos sea lion (Zalophus wollebaeki), sleeping together, likely to be related, Santa Fe Island, Galapagos, South America