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Bengal Tiger (Panthera tigris) Arrowhead patrolling his territory, Ranthambore National Park, India
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at baixa do meros, a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores
Wild Mustang stallion chasing another away from his family in White Mountain Herd Area, Wyoming, USA. August
White rhinoceros (Ceratotherium simum) two males fighting, in evening light, Sabi Sand Game Reserve, South Africa
Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Two wild Namib stallions fighting, Namib Nakluft National Park, Namib Desert, Namibia, September 2008
Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Territorial Rufous-tailed hummingbird (Amazilia tzacatl) guarding Heliconia in bloom (Heliconia champneiana) La Selva Station, Costa Rica
RF- Grey Wolf (Canis lupus) howling at sunset (digitally enhanced)
Red foxes (Vulpes vulpes) fighting in snowy field with woodland in the background, Vosges, France, February
Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
Mute swan (Cygnus olor) territorial adult, aggressive display posture, Derbyshire, UK, March
Silhouette of Eastern Grey Kangaroo {Macropus giganteus} males boxing, New South Wales, Australia
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
