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Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Atlantic grey seal (Halichoerus grypus) resting vertical beneath the surface, Lundy Island, Devon, UK. July
A young Grey Seal (Halichoerus grypus) above kelp in the Farne Islands. This seal has an orange snout from foraging on a rusty ship wreck. Northumberland, England, UK. North Sea. February
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea octopus (Benthoctopus johnsoniana) Mid-Atlantic Ridge, North Atlantic Ocean
Fangtooth (Anoplogaster cornuta) portrait, Mid-Atlantic Ridge, North Atlantic Ocean
Underwater landscape in Aare river, Canton of Berne, Switzerland, January
Giant Kelp (Macrocystis pyrifera) close up with detail of floats. Catalina Island, California, USA July
Green Hydra (Hydra viridis) walking. Digital Composite showing five movement stages. UK
Loggerhead turtle (Caretta caretta) with a shoal of Pilot fish (Naucrates ductor) Pico, Azores, Portugal, June 2009 WWE BOOK. WWE OUTDOOR EXHIBITION
Humpback whale (Megaptera novaeangliae) tail fluke above water before diving, Senja, Troms County, Norway, Scandinavia, January
Deepsea planktonic stage of crab development, Trondheimsfjord, North Atlantic Ocean
Deepsea benthic Sea cucumber (Peniagone sp) with commensal polynoid polychaete, from mid atlantic ridge, depth approx 2600m, June 2010
Deepsea benthic Sea cucumber (Peniagone porcella) from mid atlantic ridge, depth approx 2500m, June 2010
Deepsea squid from midwater catch 195-498m, Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea squid (Histioteuthis sp) from between 188m / 617ft and 507m / 1, 663ft depth, nightDeepsea squid (Histioteuthis sp) from between 188m/617ft and 507m/1, 663ft depth, night, Mid-Atlantic Ridge, North Atlantic Ocean
Octopod {Grimpoteuthis discoveryi} Mid-Atlantic Ridge, North Atlantic Ocean
Shelled Pteropod / Sea butterfly {Diacria trispinosa} with egg string attached, from between 188 and 507m, Mid-Atlantic Ridge, North Atlantic Ocean
Midwater pteropod molluscs / Sea butterflies (Clione limacina) Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea Medusa jellyfish (Colobonema sp), Mid-Atlantic Ridge, North Atlantic Ocean
Heteropod mollusc (Atlanta peroni) from between 195-498m / 640-1634ft, Mid-Atlantic Ridge, North Atlantic Ocean
Portrait of Fangtooth {Anoplogaster cornuta} Atlantic
Deepsea Anthomedusa (Pandea rubra) Gulf of Maine, Atlantic
California sea lion (Zalophus californianus) looking down from the surface. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean
Garibaldi fish (Hypsypops rubicundus) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphere Reserve, Baja California, Mexico, May
Californian sea lion (Zalophus californianus) in Giant kelp (Macrocystis pyrifera) forest, San Benitos Islands, Baja California Pacific Islands Biosphere Reserve, Baja California, Mexico, May
Humpback whale (Megaptera novaeangliae) surfacing, rostrum above surface, Bay of Fundy, New Brunswick, Canada, November
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
Orca whale (Orcinus orca) breaching. British Columbia, Canada, Pacific Ocean
Aerial view of Humpback whales (Megaptera novaeangliae) Bay of Fundy, Canada. November
Underwater view of Waterlilies (Nymphaea alba) in a lake. Alps, Ain, France, June
Sea otter (Enhydra lutris) female and pup playing. Alaska, USA, February
Sunbeams, refracted on a shallow seabed. Alghero, Sardinia, Italy. Mediterranean Sea
Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp)Pollack (Pollachius pollachius) juveniles, swimming amongst Red pom-pom seaweeds (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll, Inner Hebrides, Scotland, UK
Blue lion's mane jellyfish (Cyanea lamarckii) drifting through a seaweed forest with Red pom-pom seaweed (Ceramium sp.) and Spaghetti seaweed (Himanthalia elongata), Island of Coll
Split level photo of Sockeye salmon (Oncorhynchus nerka) swimming upstream, migrating back to the river of its birth to spawn. Trees showing autumnal colours
Great scallop (Pecten maximus) eyes, close up, Ronas Voe, Shetland, Scotland, North Atlantic Ocean, UK
Jewel anemones (Corynactis viridis), green form, close up, Bressay, Shetland, Scotland, North Atlantic Ocean, UK
Dead-mans finger (Alcyonium digitatum), close up of feeding polyps, Levenwick, Shetland, Scotland, North Atlantic Ocean, UK
Common hermit crab (Pagurus bernhardus) with Snail fur (Hydractinia echinata) growing on its shell, close up, Ronas Voe, Shetland, Scotland, North Atlantic Ocean, UK
A bloom of Sea goosberries (Pleurobrachia pileus), Birsay, Shetland, Scotland, North Atlantic Ocean, UK
Young and adult Shags (Phalacrocorax aristotelis) gathered on rocks in the sea. Treshnish Isles, Southern Hebrides, Scotland, UK. Sea of Hebrides, North East Atlantic Ocean
