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Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Red throated diver (Gavia stellata) on lochan in dawn mist, Cairngorms National Park, Scotland, UK, May
Jewel anemone (Corynactis viridis), North Uist, Outer Hebrides, Scotland, UK, May. Small repro only
White waterlily (Nymphaea alba) in a peat bog lake, De Weerribben-Wieden National Park, Holland. September
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Sperm whale (Physeter macrocephalus) tail fluke above water during dive, Kaikoura, New Zealand, July, Vulnerable species
Blue sharks (Prionace glauca) Faial Island, Azores, Portugal, Atlantic Ocean, September
Common dolphins (Delphinus delphis) pair in mating ritual, Atlantic ocean, Portugal, July
Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
RF - Bottlenose dolphins (Tursiops truncatus) porpoising, Chanonry Point, Moray Firth, Highlands, Scotland. August
Bottlenose dolphin (Tursiops truncatus) porpoising, Moray Firth, Highlands, Scotland, UK. June
European river otter (Lutra lutra) swimming in the sea, Isle of Mull, Inner Hebrides, Scotland, UK, December
European river otter (Lutra lutra) portrait, in river, Dorset, UK, November
Silhouette of Common toad (Bufo bufo) swimming in River Orchy, Highlands, Scotland, UK, April
Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Pink shrimp (Pandalus montagui), Loch Carron, Ross and Cromarty, Scotland, UK, April
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Large bull (male) Grey seal (Halichoerus grypus) chewing on a blade of kelp, resting between boulders, off Lundy Island, Devon, UK, June
Tassled anglerfish (Rhycherus filamentosus) is almost perfectly camouflaged as it hides amongst marine life, waiting to ambush prey attracted to worm life lure
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
Spotted handfish (Brachionichthys hirsutus) portrait, Derwent River estuary, Hobart, Tasmania, Australia
Afternoon sunrays dancing over rocks, in shallow water. Capo Comino, Sardinia, Italy. Mediterranean Sea
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
An Australian Sea Lion (Neophoca cinerea) bathing in the sun at the surface. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Draughtsboard Shark / Australian Swellshark (Cephaloscyllium laticeps) swims through a giant kelp forest (Macrocystis pyrifera)
Yellow cluster anemone (Parazoanthus axinellae) surrounded by Encrusting sponge (Spirastrella cunctatrix) San Pietro Island, Sardinia, Italy, Mediterranean, July
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait
A tiny amphipod (Iphimedia obesa) living on Dead mans fingers soft coral (Alcyonium digitatum) Loch Carron, Ross and Cromarty, Scotland, UK, April
Light Bulb Sea Squirt (Clavelina lepadiformis). Channel Islands, UK, June
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Hydromedusa / Trachymedusa (Crossota sp) from approx 2700m, mid Atlantic ridge, June 2010
Southern right whale (Eubalaena australis) with calluses covered in parasitic crustaceans named cyamids or whale lice (Cyamus ovalis)
Grey seal (Halichoerus grypus) laughing, Helgoland, Germany
Tail of Southern right whale (Eubalaena australis) at sunset, Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean, October
Young Grey seal (Halichoerus grypus) exploring a seaweed garden in summer, Cairns Of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, June
Old male Grey seal (Halichoerus grypus) with mouth wide open showing worn teeth, Lundy Island, Bristol Channel, England, UK, May. 2020VISION Book Plate
Blue shark (Prionace glauca) viewed from below, with Snells window effect, Pico Island, Azores, Portugal, Atlantic Ocean
Silhouette of Southern right whale (Eubalaena australis) Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean, October
Tench (Tinca tinca) on the surface of a turbid marsh pond, with Rudd (Scardinius erythrophthalmus) above, Fribourg, Switzerland, 2004
Looking down at Chubb (Leuciscus cephalus) at surface with reflection of bridge, Rhine River, Switzerland, 1993. No release available
Kangaroo Ridge in the North Cascades, in winter, Okanogan National Forest, Washington, USA, November 2010
Blue Shark (Prionace glauca) near sea surface and divers in background. Santa Maria, Azores, September
Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
