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Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Sperm whale (Physeter macrocephalus) resting just beneath surface, Faial Island, Azores, Portugal, Atlantic Ocean July
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Crab spider (Thomisus onustus) Sierra de Grazalema Natural Park, southern Spain. May
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Blue butterfly (Lycaenidae) frozen in ice, Cortes de la Frontera, Los Alcornocales Natural Park. Spain
Red spider mite (Tetranychus urticae), Andalusia, Spain, June
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
RF - Head and mane of grey Andalusian stallion running in bullfighting arena, Southern Spain, Europe
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Aquatic plants in a spring of the Gacka River, Croatia. October 2012. Photographed for The Freshwater Project
Regal jumping spider (Phidippus regius) captive male with iridescent fangs. Italy
Golden green stag beetle (Lamprima sp. ), adult male with big mandibles and a shinyGolden green stag beetle (Lamprima sp.), adult male with big mandibles and a shiny iridescent coloration, Italy. Captive
Pearlfish (Carapus acus) in anus of Sea cucumber (Stichopus regalis). These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean
Chamois (Rupicapra rupicapra) running down ridge, in early morning light. Lausons Valley, Gran Paradiso National Park, Italy, September
Fire salamander (Salamandra salamandra) female almost ready to give birth to her larvae into the stream, Apennines. Antola Regional Park, Italy
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August
Cave beetle (Leptodirus hochenwartii) a true troglobite (living only in caves). Endemic to certain karstic caves in the western Dinaric Alps. Slovenia, April
Olm (Proteus anguinus) close up of front limb, these only have three toes whilst the rear limbs have only two. Captive, Slovenia
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Spring Crocus (Crocus vernus) in flower in snow, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May
Eagle fern (Pteridium aquilinum) frond unfurling, Montseny Natural Park, a UNESCO National Park and Biosphere Reserve, Barcelona, Catalonia, Spain, June 2016
RF - Marbled white butterfly (Melanargia galathea) on knapweed, with soft focus leaves, Aosta Valley, Gran Paradiso National Park, Italy
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
Stupiste cape, Vis Island, Croatia, Adriatic Sea, Mediterranean
Canopy of a Beech woodland (Fagus sylvatica) in autumn through a circular fisheye lens. Plitvice Lakes National Park, Croatia. November
Sau and Vilanova de Sau, viewed from Tavertet and Santa Maria del Corco, Les Guilleries Natural Park, Barcelona, Catalonia, June 2016
Evening light on Gruppo del Sorapiss, Dolomite Mountains, Belluno Province, Veneto, Italy, September 2015
Lago Bain de Dones with Tofana de Rozes and Torri del Falzarego reflected, Dolomite Mountains, Belluno Province, Veneto, Italy, September 2015
Lago di Landro at dawn, Dolomite Mountains, Sud Tirol / Alto Adige, ItalyLago di Landro at dawn, Dolomite Mountains, Sud Tirol/Alto Adige, Italy
Heart-shaped leaf of Alpine Cyclamen (Cyclamen purpurascens). Plitvice Lakes National Park, Croatia. January
Spring crocus (Crocus vernus) flowering en masse on the Campo Imperatore, Abruzzo, Italy. April
Opencast mine, Rio Tinto - Red River, Sierra Morena, Gulf of Cdiz, Huelva, Andalucia, Spain. January
Crystal clear, shallow seas, Porto Pino, Sardinia, Mediterranean Sea
Iberian ibex (Capra pyrenaica), adult male, walking along mountain ridge. Sierra Nevada National Park, Andalusia, Spain. June
Iberian lynx (lynx pardinus) in distance, silhouetted on horizon, Sierra de Andujar NP, Andalusia, Spain. December
Alpine landscape with snow covered trees and clouds, Gran Paradiso National Park, Italy, March
Alpine ibex (Capra ibex) male in deep snow on a ridge during heavy snowfall, Gran Paradiso National Park, the Alps, Italy
RF - European goldfinch (Carduelis carduelis) perched in field of Sunflowers (Helianthus annuus), Cadiz, Andalusia, Spain. July. (This image may be licensed either as rights managed or royalty free.)
Valley with autumnal forest in the Estrela Mountain Range, Portugal, November 2014
Beech tree (Fagus sylvatica), Gorbeia Natural Park, Alava, Basque Country, Spain, May 2015
Sand ripples on seabed and sun burst in shallow, clear water. Capo Comino, Sardinia, Mediterranean Sea
Sunbeams, refracted on a shallow seabed. Alghero, Sardinia, Italy. Mediterranean Sea
Two Iberian ibex (Capra pyrenaica) walking along mountain ridge. Sierra Magina National Park, Andalusia, Spain. October
