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Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp)Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp.) Gam Island, Raja Ampat Islands, West Papua, Indonesia
Komodo dragon (Varanus komodoensis) male, foraging in mangove / inter-tidal zone at low tide, Komodo Island, Komodo National Park, Indonesia. Endangered
Two Komodo dragons (Varanus komodoensis) males, fighting on a remote beach, Komodo Island, Komodo National Park, Indonesia. Endangered
Flower (Barringtonia sp. ) fallen on black lava sand beach, Tangkoko National Park, northern Sulawesi, IndonesiaFlower (Barringtonia sp.) fallen on black lava sand beach, Tangkoko National Park, northern Sulawesi, Indonesia
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Ocellated eagle ray (Aetobatus ocellatus) searching for food in the sand over the seafloor, Mu Koh Similan National Park, Andaman Sea, Phang-nga, Thailand
Large school of juvenile Fusilier (Caesionidae) fish swimming in water column, Andaman Sea, Mu Koh Similan National Park, Phang-nga, Thailand
Japanese pinecone fish (Monocentris japonica) under a ledge in the reef alongside a Giant moray eel (Gymnothorax javanicus) and Banded coral shrimps (Stenopus hispidus), Richelieu Rock
White-eyed moray (Muraena thyrsoides) close up head portrait, Bali, Indonesia, Indo-Pacific
Fimbriated moray / Darkspotted moray (Gymnothorax fimbriatus) peering out from crevice with two Banded coral shrimp (Stenopus hispidus) close by, Philippines, Pacific Ocean
A venomous Banded sea krait / Yellow-lipped sea krait (Laticauda colubrina) swimming over a coral reef, Nusa Penida, Bali, Indonesia, Pacific Ocean
Whale shark (Rhincodon typus), an endangered species, feeding on zooplankton in nutrient-rich murky water, being followed by a school of Cobia (Rachycentron canadum), Adang-Rawi Archipelago, Satun
Large school of Bigeye trevally (Caranx sexfasciatus) swimming midwater, Adang-Rawi Archipelago, Andaman Sea, Satun, Thailand
Tawny nurse shark (Nebrius ferrugineus) resting in a small cave during the day, Mu Koh Similan National Park, Andaman Sea, Phang-nga, Thailand
Decomposing carcass of a juvenile Blacktip reef shark (Carcharhinus melanopterus) caught / entangled in discarded ghost nets, Phuket, Thailand, Andaman Sea
Bobtail squid (Heteroteuthis hawaiiensis) on the seabed, Komodo National Park, Indonesia, Pacific Ocean
Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp)Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp.), Kubah National Park, Sarawak, Borneo
RF - Eschmeyers scorpionfish (Rhinopias eschmeyeri) composite image showing different colour variations on black background. Lembeh Strait, North Sulawesi, Indonesia
RF - Broadclub cuttlefish (Sepia latimanus) composite image showing variation of colour and shape, on black background, Lembeh Strait, North Sulawesi, Indonesia
Mimic leatherjacket / Blacksaddle mimic filefish (Paraluteres prionurus) adult and juvenile, composite image on black background, Bali, Indonesia, Indo-Pacific
Honeycomb / Cushion starfish (Pentaceraster alveolatus) composite image on black background showing colour variations Malapascua Island, Philippines, Indo-Pacific species
Close up of hands of a Bornean Orangutan (Pongo pygmaeus), Tanjung Puting National Park, Borneo
Orange backed red langur (Presbytis rubicunda chrysea) in rainforest understorey. Danum Valley, Sabah, Borneo, Malaysia
Infant Bornean Orang-Utan (Pongo pygmaeus) clinging to its mother. Camp Leakey, Tanjung Puting NP, Kalimantan, Borneo
Western tarsier (Tarsius bancanus) clinging to tree, Danum Valley, Sabah, Borneo
Venomous Yellow-lipped sea krait (Laticauda colubrina) heading up to the surface to breathe, Philippines
Blackspotted puffer (Arothron nigropunctatus) curled up on the reef for the night, with a transparent cleaner shrimp (Urocaridella antonbrunii) inspecting its tail, Philippines
Smoke billowing from Mount Bromo volcano. View from the summit of Mount Batok. Bromo Tengger Semeru National Park, Java, Indonesia, July, 2013
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
A massive school of millions of sardines (Sardina pilchardus), Moalboal, Cebu, Central Visayas, Philippines
Ornate ghost pipefish (Solenostomus paradoxus), juvenile, hides within a crinoid, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Montane horned frog (Megophrys kobayashii) head close up, Kinabalu National Park, Sabah, Borneo, Malaysia
Portrait of a lionfish (Pterois volitans), a study of concentration as it hunts. Salawati Island, Raja Ampat, Indonesia. Ceram Sea
Minute / Dwarf filefish (Rudarius minutus) just over an inch long as an adult, Celebes Sea, Mabul Island, Malaysia
Black crested monkey or macaque (Macaca nigra) infant in tree, Sulawesi, Indonesia. Critically endangered
Bromo volcano, Java, Indonesia
Pair of Coleman shrimps (Periclimenes colemani) living in Fire urchin (Asthenosoma varium). The female is the larger shrimp. Anilao, Batangas marine protected area, Luzon, Philippines
Wide angle macro photo of an algae octopus (Abdopus aculeatus) as it reaches out and investigates the camera. Dauin, Dauin Marine Protected Area, Dumaguete, Negros, Philippines
Sunda leopard cat (Prionailurus javanensis) kitten. Sabah, Malaysian Borneo
Blyths paradise flycatcher (Terpsiphone affinis) male calling, Sabah, Malaysian Borneo
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Elegant conch (Lentigo pipus) close up of eye stalks, Philippines
Sumatran tiger (Panthera tigris sondaica) portrait, captive
Lowland dipterocarp rainforest, with huge Tualang (Tapang) or Mengaris Tree (Koompassia excelsa) in the foreground, Danum Valley, Sabah, Borneo
Western / Horsfields tarsier (Tarsius bancanus) hunting invertebrate prey in rainforest understory vegetation at night. Danum Valley, Sabah, Borneo
Giant shield bug nymphs (Pycanum sp. ), recently hatched from eggsGiant shield bug nymphs (Pycanum sp.), recently hatched from eggs. Danum Valley, Sabah, Borneo
Safety in Numbers - Giant Shield bug nymphs (Pycanum sp.), recently hatched from eggs, with a final individual still to emerge