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Elephant (Loxodonta africana) female and calf under tree in the rain, Masai-Mara game reserve, Kenya
RF- Whooper Swan (Cygnus cygnus) standing on frozen loch at dawn in winter. Loch Insh, Cairngorms National Park, Scotland, UK
Poppy bee (Osmia papaveris) resting on Common poppy (Papaver rhoeas) cutting piece of petal to use as nesting material, Bamberg, Germany. June
Red fox (Vulpes vulpes) vixen walking through snow, Derbyshire, UK. January
Puma (Puma concolor) walking with the Torres del Paine mountains in background, Torres del Paine National Park, Magallanes, Chile
Humpback whale (Megaptera novaeangliae) diving into the depths, Hawaii, Pacific Ocean
Single almond tree (Amygdalus communis), with wind turbines in background. Sisante, Albacete, Spain
Roe deer (Capreolus capreolus) fawn, Vosges, France, June
RF - Wandering albatross (Diomedea exulans) flying over the rugged coast of South Georgia. South Atlantic. (digitally stitched image)
Red fox (Vulpes vulpes) cub exploring on fallen tree in woodland, Switzerland
Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019
Dusky Shark (Carcharhinus obscurus) solitary shark patrols the deep blue waters off Port St Johns, South Africa
Reticulated Giraffe (Giraffa camelopardalis reticulata) standing in grassland. Laikipia, Kenya. October
RF- New Forest pony near Bratley View, New Forest, Hampshire, UK. November
Wolf (Canis lupus) lone male walking in shadow of the Teton Mountains. Snake River valley near Jackson Hole. Grand Teton National Park, Wyoming, USA. January
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
English oak tree (Quercus robur) in field of Oilseed rape / rapeseed (Brassica napus), UK, May 2017
Black and white photograph of saltmarsh at Annat, Beinn Torridon, Scotland, UK, November
Blacktip reef shark (Carcharhinus melanopterus) just at the surface off the island of Yap, Micronesia
Thonga Beach on the coast of Indian Ocean, Maputuland, KwaZulu-Natal, South Africa, January
Whooper swan (Cygnus cygnus) standing alone in snow storm, Hokkaido Japan February
RF- Arabian Oryx (Oryx leucoryx) Dubai Desert Conservation Reserve, Dubai, UAE
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
Emperor penguin (Aptenodytes forsteri) walking, Gould Bay, Weddell Sea, Antarctica
RF - Strawberry Poison Frog (Oophaga pumilio). Central Caribbean foothills, Costa Rica
RF - Coyote (Canis latrans) in snow, Yellowstone. February
Coyote (Canis latrans) in wintry landscape, Yellowstone National Park, USA, February
Bighorn Sheep (Ovis canadensis) on the hillside, ruminating, Yellowstone National Park, Wyoming, USA. January 2011
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
RF- Bison (Bison bison) standing in steam from geothermal springs. Yellowstone National Park, Wyoming, USA, February
A lone hiker stands against blizzard conditions on An Teallach. Ullapool, Highlands of Scotland, UK, January 2016
Male black rhinoceros (Diceros bicornis) silhouetted against clouds, Masai-Mara Game Reserve, Kenya. March
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Capercaillie (Tetrao urogallus) in forest habitat, Jalasjarvi, Finland, April
African Blue tit (Cyanistes teneriffae) Morocco, February
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Young Baikal seal (Pusa sibirica) at breathing hole. Lake Baikal, Russia, April
Sycamore tree (Acer pseudoplatanus), growing on limestone pavement, Lancashire, UK. May
Fin whale (Balaenoptera physalus) surfacing with mountain landscape, Liefdefjorden, Svalbard, Norway. July 2011
Red lechwe (Kobus leche) silhouetted at sunrise, Okavango Delta, Botswana
tail detail of Sperm Whale (Physeter macrocephalus) Pico Island, Azores, Portugal, Atlantic Ocean
Iberian black pig foraging in oak woodland, Sierra de Aracena Natural Park, Huelva, Andalucia, Spain, Europe. Breed used to produce Iberico ham / Jamon Iberico
Green Turtle (Chelonia mydas) female on beach, Concha Beach, municipality of Itacare, Southern Bahia State, Eastern Brazil
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, walking down a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Hanuman / Northern Plains Grey Langur (Semnopithecus entellus) youngster sitting on the trunk of a Banyan tree, Ranthambore National Park, Rajasthan, India, June
Harlequin duck (Histrionicus histrionicus) male on a large boulder in the fast flowing and turbulent waters of the Laxa River, Iceland, May. Bookplate from Danny Greens The Long Journey North
Harvest mouse (Micromys minutus) on wheat stem, Devon, UK captive
