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RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Red fox (Vulpes vulpes) silhouetted, Sado Estuary, Portugal. September
Synchronous fire flies (Pteroptyx sp. ) mangroves along the Kinabatangan River, Sabah, BorneoSynchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Praying mantis nymph silhouetted through leaf, . Raja Ampat, Western Papua, Indonesian New Guinea
Chinese lanterns (Physalis alkekengi) skeletons, silhouette on white background
Hedge parsley (Torilis japonica) close up pattern of dead seed heads. Silhouette on white background
Chinese Lanterns (Physalis alkekengi), Hydrangea flowers, Poplar leaves (Populus) and Honesty skeletons (Lunaria annua), silhouettes on white background
Ripe Barley (Hordeum vulgare) ears black and white backlit image
Stonehenge at sunrise, Salisbury Plain, Wiltshire, England, UK, June 2006
Corfe Castle in morning mist, Corfe, Isle of Purbeck, Dorset, England, UK. September 2013
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
Angolan giraffes (Giraffa camelopardalis angolensis) and black rhinoceros (Diceros bicornis) reflecting in waterhole at sunset, Okaukuejo, Etosha National Park, Namibia, Africa
Nilgai or Blue bull (Boselaphus tragocamelus), silhouette of male at sunset, with Black drongo bird on its back. Velavadar National Park, Gujarat, India
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic
Mountain landscape at Tangjiahe National Nature Reserve, Sichuan Province, China April, 2015
Ripe Barley (Hordeum vulgare) ears in field ready for harvest and the rising sun, Stone Henge, Salisbury, England, UK, August
Peruvian mantis (Oxyopsis peruviana) silhouette with sun behind, captive, occurs in South America
Groot Schietveld at sunrise, Wuustwezel, Belgium, May 2010
Scots pine trees (Pinus sylvestris) on island in wetlands, Klein Schietveld, Brasschaat, Belgium
Boababs (Adansonia grandidieri) silhouetted at dawn, Allee des Baobabs / Avenue of the Baobabs, Morondave, Madagascar
Pheasant (Phasianus colchicus) male flying to roost at sunset with trees silhouetted in background England, UK. December
RF - Grizzly Bear (Ursus arctos) at dawn, Lake Clarke National Park, Alaska, September
RF- White storks (Ciconia ciconia) nesting on poles erected by the city of Caceres in Extremadura, Spain, to compensate for the removal of an old farmhouse supporting a colony of storks
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013
View from Isle of Skye across Sound of Rsay to Rona and Torridon Hills, at dawn, Inner Hebrides, Scotland, UK, April 2014
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
Wood sandpiper (Tringa glareola) at dusk silhouetted against setting sun, Karigasniemi, Northern Finland, May. Bookplate from Danny Greens The Long Journey North
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
RF - Delicate grass and flowers silhouetted black against white background
European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March