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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Sea stacks and cliffs at sunset at Eshaness / Esha Ness, peninsula in Northmavine on the island of Mainland, Shetland Islands, Scotland, UK, May 2018
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at sunrise with Slioch in background, Wester Ross, Scotland, UK, November 2014
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Super moon rising above Glastonbury Tor with people watching, Somerset, England, UK, 27th September 2015
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Knoydart and Loch Nevis from Sgurr Cos na Breachd-laoidh with sun shining through cloud, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at dawn with Slioch in background, Wester Ross, Scotland, UK, November 2014
Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Two Rooks (Corvus frugilegus) silhouetted as they perch on a tree branch at their roost site at sunset, Gloucestershire, UK, February
Chinese lanterns (Physalis alkekengi) skeletons and Honesty seed pods (Lunaria annua)
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Helicopter flying over volcanic eruption near Eyjafjallajoekull glacier, Iceland, 24th March 2010. Volcano previously dormant since 1821
Star trails above Scots pine (Pinus sylvestris) woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, March 2017
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Red fox (Vulpes vulpes) silhouetted in mixed beech-Douglas fir forest. Freiburg, Black Forest National Park, Germany, October
Sun rays shining through clouds onto Loughrigg Fell and hills of the Lake District National Park, in evening. Ambleside, Cumbria, England, UK. April 2020
Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Mongol driving a camel caravan of Bactrian camel (Camelus bactrianus) Bashang Grassland, Zhangjiakou, Hebei Province, Mongolia, China
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Silhouette of an African leopard (Panthera pardus pardus) snarling, Mkuze, South Africa. September. White Fang - this leopard bared its fangs to the hyenas threatening his prey
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Quiver trees (Aloidendron dichotomum) silhouetted against dawn sky, Namibia, May
Avielochan at sunrise, Cairngorms National Park, Scotland, UK, August
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Humpback Whale (Megaptera novaeangliae) tail fluke at surface as whale dives, in the late afternoon light, Andenes, Andoya island, North Atlantic Ocean, Norway Janurary
Teasel (Dipsacus fullonum), Hedge parsley (Torilis) and Allium seedhead silhouettes on white background
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic, volumetric lighting, Hungary, January
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
The Milky Way over Sand Beach, Acadia National Park, Maine, USA. July, 2020
Female Fox (Vulpes vulpes) walking on tree trunk in heavy snowstorm in the dark, Verteskozma, Hungary
Mute swan (Cygnus olor) swimming on sun-dappled water, Lower Silesia, Poland
Banded demoiselle (Calopteryx splendens) male silhouetted on a fern, Lower Tamar Lakes, Cornwall, UK. July
Volcanic eruption with silhouette of tree, Cumbre Vieja Volcano, La Palma, Canary Islands. September 2021
View of the Alps from the Aiguille du Midi, Mont Blanc, near Chamonix, Savoie, France
Fallow deer (Dama dama) hind and fawn in early morning mist. London, UK. September
Crested tit (Lophophanes cristatus) silhouette, Sierra de Grazalema Natural Park, Cadiz, Andalusia, Southern Spain. October
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic volumetric lighting. Vertes Mountains, Hungary
Pine (Pinus sp) trees on Colmer's Hill, in morning mist. Near Bridport, Dorset, England, UK. September 2012
Tibetan antelope / Chiru (Pantholops hodgsonii) three silhouetted, Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
European Robin (Erithacus rubecula) backlit on branch, Sierra de Grazalema Natural Park, southern Spain, October
Golden Eagle adult (Aquila chrysaetos) silhouetted in tree after sunset, Kuusamo, Finland, January
Night sky over Stone arch, called the Julia arch. Ennedi Natural and Cultural Reserve, UNESCO World Heritage Site, Chad. September 2019
Bactrian camel (Camelus bactrianus) male silhouetted at sunset, living in the wild but owned by a camel herdsman, Turpan Basin, Gobi Desert, Xinjiang, China
Huge boulder balanced on rocky outcrop at sunset, Namib desert, Namib-Naukluft National Park, Namibia, March
Red fox (Vulpes vulpes) silhouetted, running along horizon, Switzerland
Aurora Borealis above silhouetted trees, northern Finland, March
Frozen water patterns in Klein Schietveld, Brasschaat, Belgium
African elephant (Loxodonta africana) two silhouetted at sunset with goose flying overhead, Mkuze, South Africa
Goat willow (Salix caprea) in mist at dawn, full moon in sky. Klein Schietveld, Brasschaat, Belgium. August 2019
Short-toed Eagle (circaetus gallicus) flying amongst Stone Pines (Pinus pinea). Coto Donyana, Andalusia, Spain. 2007
Red fox (Vulpes Vulpes) silhouetted in door to a house, North London, England, UK. June
Silhouette of the head of a Grey-headed flying-fox (Pteropus poliocephalus) through its wings
Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China
Little penguin (Eudyptula minor) standing on rock, silhouetted by Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Beech trees (Fagus sylvatica) at night in winter, with lamp post illuminating the woodland. Felbrigg Woods, Norfolk, England, UK. January
Morning mist over Vosges mountain, France, October
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Red fox (Vulpes vulpes) silhouetted, Sado Estuary, Portugal. September
Synchronous fire flies (Pteroptyx sp)Synchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Moonlight over the Skellig Islands, Puffin Island, County Kerry, Ireland, Europe. September 2015
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Praying mantis nymph silhouetted through leaf, . Raja Ampat, Western Papua, Indonesian New Guinea
Chinese lanterns (Physalis alkekengi) skeletons, silhouette on white background