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African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Common blue damselfly (Enallagma cyathigerum) roosting among dewy grasses, Tamar Lakes, Cornwall, UK. June
RF - Common blue butterflies (Polyommatus icarus) resting on a backlit grass, evening light, Vealand Farm, Devon, UK. June
RF - Wild garlic / Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UKRF - Wild garlic/Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UK. May
Poppy (Papaver rhoeas) Groot Schietveld, Wuustwezel, Belgium, July
Demoiselle cranes (Anthropoides virgo) flock, India
RF- Mute swan (Cygnus olor) close up peering at camera, juvenile. Fife, Scotland, UK, November
Tropical Screech Owl (Megascops choliba) perched in tree hole in rainforest canopy. Cristalino State Park, Amazonia, Brazil, South America
Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Great White Egret (Egretta alba) with water splashing from beak, Hungary January
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Marbled white butterfly (Melanargia galathea) resting on reed, Devon, UK, July
Cowslips (Primula veris) growing on organic farm. Norfolk, UK, April
Two male Bearded reedling (Panurus biarmicus) eating seeds from the spike of a common bulrush (Typha latifolia) in a reedbed, Belgium, January. Digital composite
Small Yellow Underwing moth (Panemeria tenebrata) on stitchwort (Stellaria) South Karelia, southern Finland, June
Northern Hawk-Owl (Surnia ulula) seen through branches, southwest Finland, February
Scottish wildcats (Felis sylvestris), captive, UK, June
Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013
Kittens exploring garden lawn, Germany
Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Natures Best Photography Windland Smith Rice International Awards Competition
Golden-crowned Sifaka or Tattersalls Sifaka (Propithecus tattersalli) climbing down tree in forests near Andranotsimaty, Daraina, north east Madagascar. Critically Endangered
Phayers Leaf monkey (Trachypithecus phayrei) Gaoligongshan NP, Yunnan province, China
Greater Stitchwort (Stellaria holostea) and Bluebells (Hyacinthoides non-scripta) in flower, UK, May
Swallowtail (Papilio machaon) in flight, Finland, Europe
Female goshawk (Accipiter gentilis) in flight, just after taking off from perch. Southern Norway, January
Fallow deer (Dama dama) one deer stops and looks whilst the herd runs on, with sunlit leaves out of focus, Vastmanland, Sweden
Pair of Emerald damselflies (Lestes sponsa) resting on a reed, with only tips of wings in focus, Cornwall, England, UK, August
RF - Common spotted orchids (Dactylorhiza fuchsii), Volehouse nature reserve, Devon, UK, June
White stork (Ciconia ciconia) adult portrait, captive, Vogelpark Marlow, Germany, May
European wild cat (Felis silvestris) kitten portrait, Bavarian Forest National Park, Germany, captive
Gentoo penguins (Pygoscelis papua) on the shore of Deception Island, Antarctica
Japanese Macaque (Macaca fuscatata) juvenile, reaching out to touch the camera. Jigokudani, Nagano Prefecture, Honshu, Japan
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Leopard (Panthera pardus) back of head, Samburu National Reserve, Kenya
Frigga Fritillary (Boloria frigga) in habitat over cotton grass (Eriophorum) Finland, June
Marbled white butterfly (Melanargia galathea) nectaring on flower, Vallee de la Seille (Seille Valley), Lorraine, France, June
Apollo butterfly (Parnasius apollo) feeding on flower, Mount Terminillo, Rieti, Lazio, Italy, July 2011
Spotted lancehead (Bothrops punctatus) with tongue extended curled round branch, Canande, Esmeraldas, Ecuador
Puyango whorltail iguana (Stenocercus puyango) portrait, Macara, Loja, Ecuador
Male Choco trogon (Trogon comptus) perched on branch, Mashpi, Pichincha, Ecuador
Tambococha rain frog (Pristimantis omeviridis) artistic view of eye, Yasuni National Park, Orellana, Ecuador
Ling heather (Calluna vulgaris) Peak District National Park, Derbyshire, UK, August
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
Ling Heather (Calluna vulgaris) with sun setting behind, Peak District National Park, Derbyshire, UK. August
Dalmatian Pelicans (Pelecanus crispus) portrait. Lake Kerkini, Greece, March 2012
Common poppies (Papaver rhoeas) backlit in evening light, Crantock, Newquay, Cornwall, UK. June 2015
