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Two adult Tufted titmice (Baeolophus bicolor) surrounded by snow, winter, New York, USA
European badger (Meles meles) in woodland in late evening light. Scotland, UK, August
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Oxeye daisies (Leucanthemum vulgare)in upland hay meadow, Northumberland National Park, UK, June
Little owl (Athene noctua) perched on branch of oak tree, London, UK, June
RF - Downy birch (Betula pubescens) changing to autumn colours. Caledonian pine forest, Glen Strathfarrar, Scottish Highlands. Scotland. October
Oxeye daisies (Leucanthemum vulgare) in upland hay meadow, Northumberland National Park, UK, June
Espanola lava lizard (Microlophus delanonis) female with Waved albatross (Phoebastria irrorata) behind. Punta Suarez, Espanola Island, Galapagos
Herb robert (Geranium robertianum) flower, Broxwater, Cornwall, UK. April
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
RF - Marbled white butterfly (Melanargia galathea) on knapweed with soft focus flowers, Aosta Valley, Gran Paradiso National Park, Italy
Autumn leaves moving in the wind, Plitvice Lakes National Park, UNESCO World Heritage Site, Central Croatia. Croatia
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
RF - Common darter dragonfly (Sympetrum striolatum) resting on reed, dew covered, surrounded by webs, Vealand, Devon, UK. August
Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Globe Thistle (Echinops ritro), dead in winter landscape with Alpine ibex (Capra ibex) silhouet behind, Gran Paradiso national park, Italian Alps, Italy, January
Katydid (Tettigoniidae) with Giraffe (Giraffa camelopardalis) in the background, Masai Mara National Park, Kenya
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Great egret (Ardea alba) group of five at waters edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February
RF - Eschers blue butterfly (Polyommatus escheri) on grass, with soft focus blue flowers, Aosta Valley, Gran Paradiso National Park, Italy
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Hoopoe (Upupa epops) with European roller (Coracias garrulus) in the background. Pusztaszer, Hungary. May
Dew on spiderweb with Common poppy (Papaver rhoeas) in the background, Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Dew on spiderweb with Grey-leaved cistus (Cistus albidus) Sierra de Grazalema Natural Park, El Bosque, southern Spain, April
Orchid (Orchis langei) in sunset light, Sierra de Grazalema Natural Park, southern Spain, May
Orchid (Orchis olbiensis), Sierra de Grazalema Natural Park, Benaocaz, southern Spain, May
Lion (Panthera leo) portrait of two lionesses, captive, occur in Africa
African lioness (Panthera leo) sitting patiently in the long grass, watching herd of Common eland (Tragelaphus oryx). Focus on lion. Grumeti Reserve, Northern Tanzania
Common blue damselfly (Enallagma cyathigerum) roosting among dewy grasses, Tamar Lakes, Cornwall, UK. June
RF - Common blue butterflies (Polyommatus icarus) resting on a backlit grass, evening light, Vealand Farm, Devon, UK. June
RF - Wild garlic / Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UKRF - Wild garlic/Ramson flowerts (Allium ursinum) Valency Valley, Boscastle, Cornwall, UK. May
Poppy (Papaver rhoeas) Groot Schietveld, Wuustwezel, Belgium, July
Demoiselle cranes (Anthropoides virgo) flock, India
RF- Mute swan (Cygnus olor) close up peering at camera, juvenile. Fife, Scotland, UK, November
Tropical Screech Owl (Megascops choliba) perched in tree hole in rainforest canopy. Cristalino State Park, Amazonia, Brazil, South America
Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Great White Egret (Egretta alba) with water splashing from beak, Hungary January
Feral domestic rabbit (Oryctolagus cuniculus) standing on hind legs on coast, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Marbled white butterfly (Melanargia galathea) resting on reed, Devon, UK, July
Cowslips (Primula veris) growing on organic farm. Norfolk, UK, April
Two male Bearded reedling (Panurus biarmicus) eating seeds from the spike of a common bulrush (Typha latifolia) in a reedbed, Belgium, January. Digital composite
Small Yellow Underwing moth (Panemeria tenebrata) on stitchwort (Stellaria) South Karelia, southern Finland, June
Northern Hawk-Owl (Surnia ulula) seen through branches, southwest Finland, February
Scottish wildcats (Felis sylvestris), captive, UK, June
Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013
