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Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March
RF - Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean
RF - Seafans (Melithaea sp. ) growing on dropoff, beneath schooling fish. Boo WindowsRF - Seafans (Melithaea sp.) growing on dropoff, beneath schooling fish. Boo Windows, Boo Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
Bridled parrotfish (Scarus frenatus) clownish grin reveals its power tools: grinding teeth used to scrape algae from rock, Maldives, Indian Ocean
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, with scuba diver in background, Arctic circle Dive Center, White Sea, Karelia, northern Russia, captive
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Detail of magnificent Sea anemone (Heteractis magnifica) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Blue Whales (Balaenoptera musculus) throat inflated, filter feeding at sea surface. California, USA. Sequence 1 of 2
Humpback Whale (Megaptera novaeangliae) calf, Tonga, South Pacific, September
Humpback Whale (Megaptera novaeangliae) mother and calf. Tonga, South Pacific, September
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Big fin reef squid (Sepioteuthis lessoniana) descends from open water to the reef at night. Arafur Sea, Triton Bay, West Papua, Indonesia
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Aggregation / bloom / swarm of Moon Jellyfish (Aurelia aurita) in the clear waters of Loch Na Keal, off the Isle of Mull, north west Scotland, UK, June, Photographed through the surface
Sperm Whale (Physeter macrocephalus) calf swimming uder its mother. Sao Miguel Island, Azores, July
Manta Ray (Manta Birostris) and Devil Ray (Mobula tarapacana) in background, accompanied by Pilot Fish (Naucrates ductor) and Remora. Santa Maria Island, Azores, September
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Great Blue Shark (Prionace glauca), dorsal view against dark water. Over 20 million Blue Sharks are fished annually, as bycatch and due to the high value of their fins
A portrait of a Leafy Seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Two Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp. ). Digital compositeTwo Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp.). Digital composite. Misool, Raja Ampat, West Papua, Indonesia, November
A Pygmy Seahorse (Hippocampus bargibanti) camouflaged in red Seafan (Muricella sp.). Pygmy Seahorses are small, most less than 15mm in length. Tulamben, Bali, Indonesia, October
A juvenile Weedy Seadragon (Phyllopteryx taeniolatus) in its habitat. Flinders Jetty, Melbourne, Victoria, Australia, March
A young Spiny / Yellow / Longsnouted Seahorse (Hippocampus guttulatus), in an aquarium attached to plastic seagrass leaves. This seahorse is part of a UK seahorse breeding program
Reef with Pygmy Sweepers (Parapriacanthus ransonetti), Barrel Sponge (Xestospongia testudinaria), and soft corals. Komodo National Park, Indonesia, October
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)
Group of South American fur seals (Arctocephalus australis) swimming near the rocky shore. Puerto Deseado, Patagonia, Argentina, March 2008
South African fur seal (Arctocephalus pusillus pusillus) swimming near surface, Seal Island. False Bay, South Africa, July 2010
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
California sea lion pup (Zalophus californianus) portrait in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Atlantic grey seal (Halichoerus grypus) swimming beneath the surface, Lundy Island, Devon, England, UK. July 2010. Highly commended, Coast and Marine category
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas
Bottlenose dolphins (Tursiops truncatus) in shallow water over a sand bank. Sandy Ridge, Little Bahama Bank, Bahamas. Tropical West Atlantic
As soon as it entered the water the young Olive ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the coast
Golden damselfish (Amblyglyphidodon aureus) swimming past coral reef wall with colourful soft corals. Bali, Indonesia
Large group of Californian sealions (Zalophus californianus) swimming underwater, Anacapa Island, California, Pacific Ocean
Bigscale soldierfish (Myripristis berndti), Sweetlips, Philippine chromis with Blueline snappers in background. Komodo National Park, Indonesia
Green turtle (Chelonia mydas) swimming over sea grass, Grenadines, Caribbean, February 2010