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Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
A young Grey Seal (Halichoerus grypus) above kelp in the Farne Islands. This seal has an orange snout from foraging on a rusty ship wreck. Northumberland, England, UK. North Sea. February
A school of Southern stingrays (Hypanus americanus) on a sand bar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Long exposure image of Tiger shark (Galeocerdo cuvier) chasing bait, Bahamas
Split level image of Caribbean reef shark (Carcharhinus perezi) breaking surface at dusk. Grand Bahama, Bahamas. Digitally manipulated image
A lone Caribbean reef shark (Carcharhinus perezi) cruises over sand ripples. Walkers Cay, Northern Bahama Islands, Republic of Bahamas
Free diver swimming amongst Californian sealions (Zalophus californianus) Anacapa Island, California, Pacific Ocean Model released
Deepsea octopus (Benthoctopus johnsoniana) Mid-Atlantic Ridge, North Atlantic Ocean
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Deepsea planktonic stage of crab development, Trondheimsfjord, North Atlantic Ocean
Deepsea squid (Histioteuthis sp) from between 188m / 617ft and 507m / 1, 663ft depth, nightDeepsea squid (Histioteuthis sp) from between 188m/617ft and 507m/1, 663ft depth, night, Mid-Atlantic Ridge, North Atlantic Ocean
Deepsea Anthomedusa (Pandea rubra) Gulf of Maine, Atlantic
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
Ornate ghost pipefish (Solenostomus paradoxus) young hides amongst the branches of a black coral bush (Antipathes sp) Sangeang Island, Sumbawa, Indonesia, Flores Sea
Illustration of Giant Octopus (Octopus dofleini) attacking a ship, 1815 copperplate engraving from Bertuchs Bilderbuch fur kinder 16 plate 5. with contemporary colour
Close-up of scales of the Yellowbar Angelfish (Pomacanthus maculosus). Red Sea, Egypt
Oriental sweetlips (Plectorhinchus vittatus), Maldives, Indian Ocean
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Pink anemonefish (Amphiprion perideraion) with anemone showing the effects of bleaching. The bleached anemone is almost white
Galapagos garden eels (Heteroconger / Taenioconger klausewitzi) on seabed, Galapagos Islands
John dory (Zeus faber) portrait, Babbacombe Bay, Devon, UK, October
Silvertip sharks (Carcharhinus albimarginatus) in open water, Revillagigedo islands, Mexico. Pacific Ocean
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Close up of tiny (10mm) Pygmy seahorse (Hippocampus bargibanti) living disguised in Muricella sp. sea fan. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Female Denises pygmy seahorse (Hippocampus denise) on coral seafan (Annella mollis) Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Curimbata (Prochilodus lineatus) in a clear stream of the Pantanal, with Water hyacinth (Eichhornia sp.), Mato Grosso do Sul, Brazil. Taken for the Freshwater Project
Killer whale (Orcinus orca) tail slapping at sunset, transient race, Vancouver Island, British Columbia, Canada, July
tail detail of Sperm Whale (Physeter macrocephalus) Pico Island, Azores, Portugal, Atlantic Ocean
Bottlenose Dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal
A portrait of a Mexican hogfish (Bodianus diplotaenia). Socorro Island, Revillagigedos, Mexico. East Pacific Ocean
Red / Blotcheye soldierfish (Myripristis murdjan) and sea whips coral, Junceella at Fotteyo caves, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in sea-anemone, Red Sea
Barrel Jellyfish (Rhizostoma pulmo) against crepuscular light rays. Sark, British Channel Islands, July
Humpback Whale (Megaptera novaeangliae) calf. Tonga, South Pacific, September
Humpback Whale (Megaptera novaeangliae) female. Tonga, South Pacific, September
Golden damselfish (Amblyglyphidodon aureus) hiding amongst the branches of red whip coral (Ctenocella sp.) Misool, Raja Ampat, West Papua, Indonesia
Portrait of Red Sea anemonefish (Amphiprion bicinctus) against open water. Nuweiba, Sinai, Egypt
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
RF - Emperor shrimp (Periclimenes imperator) exploring alien-looking landscape of host sea cucumber (Bohadschia argus). Emperor shrimps are commensals on many hosts
A female Spiny Seahorse (Hippocampus guttulatus) in weeds. Sardina, Gran Canaria, Canary Islands, East Atlantic Ocean, October
A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
In-camera motion blur image of a female Yellow Seahorse (Hippocampus kuda). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March
A batch of Leafy Seadragon (Phycodurus eques) eggs on the tail of a male. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March