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A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Dolphin trainer interacting with Bottlenose Dolphin (Tursiops truncatus), Dolphin Reef, Eilat, Israel - Red Sea. Model released
Caribbean reef squid (Sepioteuthis sepioidea) in mid water over a coral reef. East End, Grand Cayman, Cayman Islands, West Indies. Caribbean Sea
Bulb tentacle sea anemone (Entacmaea quadricolor) close up detail, Maldives, Indian Ocean
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Hydromedusa / Trachymedusa (Crossota sp) showing tentacles, from approx 2700m, mid Atlantic ridge, June 2010
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011
A Florida manatee (Trichechus manatus latirostrus) in upright posture with tail on river bed, outside Three Sisters Spring. Crystal River, Florida, USA
Stove-pipe sponge (Aplysina archeri) and Orange elephant ear sponge (Agelas clathrodes) Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Orcas / Killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Purple sea pen (Virgularia gustaviana) on sandy sea bed. Rinca, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean
Blue shark (Prionace glauca) Pico Island, Azores, Portugal, Atlantic Ocean, July
Close up of deepsea Basket star (Gorgonocephalus sp) from coral seamount, SW Indian Ridge, Indian Ocean, December 2011
Whale Shark (Rhincodon typus) accompanied by Pilot Fish (Naucrates ductor) and Remora. The biggest fish in the world has only started to visit the Azores in the past three years due to water
A silhouette of a California sealion pup (Zalophus californianus) next to cliffs. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
Lemon shark (Negaprion brevirostris) under sun rays. Little Bahama Bank, Bahamas. West Atlantic Ocean
Oceanic whitetip shark (Carcharhinus longimanus) with Pilot fish (Naucrates ductor). Tropical West Atlantic Ocean, off Cat Island, Bahamas
A large female Tiger Shark (Galeocerdo cuvier) swimming in open water. Grand Bahama, Bahamas. Tropical West Atlantic Ocean. March
Southern stingray (Hypanus americanus) swimming over sand ripples on sandbar, Stingray City, Grand Cayman. British West Indies
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Two Atlantic spotted dolphins (Stenella frontalis) over a shallow sand bank. On the left is a juvenile which does not have the characteristic spots of the adults
Large group of Sunflower sea stars (Asterias / Pycnopodia helianthoides) covering rock, pacific coast, Canada, August
Orange coloured Seashore Lichens (Xanthoria parietina) and Sea Ivory (Ramalina siliquosa) UK, August
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway
Blue jellyfish (Cyanea lamarckii), feeding on small plankton, Lundy Island Marine Conservation Zone, Devon, England, UK, May
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
