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Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Northern rockhopper penguin (Eudyptes moseleyi) parent with chick. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Emperor penguin (Aptenodytes forsteri) tobogganing, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Royal tern (Sterna maxima) perching on a sand castle on a beach, Fort De Soto Park, St. Petersburg, Florida, USA, March
Crested auklets (Aethia cristatella) pair interacting while perched on rock, St Paul Island, Pribilofs, Alaska, USA, July
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Puffins (Fratercula arctica) and Razorbills (Alca torda) with Eilean Mhuire behind, Shiant Isles, Outer Hebrides, Scotland, UK. June, 2018
King penguins 1+Aptenodytes patagonicus+1 group of three on the shore, St. Andrews Bay, South Georgia. November
King penguins 1+Aptenodytes patagonicus+1. Grytviken, South Georgia. November
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) incubating egg, Puerto Pajas, Isabela Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Northern gannet (Morus bassanus) in flight over colony, Quebec, Canada, August
Atlantic puffin (Fratercula arctica) in flight, Machias Seal Island off the coast of Maine, USA, July
Atlantic puffins (Fratercula arctica) amongst flowering Sea campion (Silene uniflora), Skomer Island, Pembrokeshire, Wales, UK. May
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Gannet (Sula bassana) diving, July UK Shetland
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Northern rockhopper penguin (Eudyptes moseleyi) portrait, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Gentoo Penguin (Pygoscelis papua) coming out of the sea, Cuverville Island, Antarctic Peninsula, Antarctica
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Emperor penguin (Aptenodytes forsteri), chick in brood pouch, Snow Hill Island, Antarctic Peninsula
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
