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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Chinstrap penguin (Pygoscelis antarcticus) with chicks, Antarctic Peninsula, Antarctica
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Emperor penguin (Aptenodytes forsteri) viewed through hole in iceberg at Snow Hill Island rookery, Antarctica. November
Black skimmer (Rynchops flavirostris) fishing by skimming along water surface, Pantanal, Brazil
Two Gentoo penguin (Pygoscelis papua) chicks sleeping huddled together, Port Lockroy, Goudier Island, Antarctic Peninsula, Antarctica
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
King penguins (Aptenodytes patagonicus) at sunrise, Falklands. Highly honoured in the Ocean View Category of the Nature's Best Windland Smith Rice Ocean View Competition 2017
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Tabular iceberg floating in Weddell Sea, iceberg broken away from Larson C ice shelf. Adelie penguin (Pygoscelis adeliae) colony dwarfed on sea ice below. Near Danger Islands, Antarctica
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
Grizzly bear (Ursus arctos horribilis) and Grey wolf (Canis lupus) on beach, Katmai National Park, Alaska, USA, August
RF- Adelie Penguin (Pygoscelis adeliae) on iceberg. Yalour Islands, Antarctic Peninsula, Antarctica. February
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Royal penguin (Eudyptes schlegeli) walks along the beach. Sandy bay, Macquarie Island, Australian Territory
Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
Puffins (Fratercula arctica) swimming underwater. Puffins spend most of their lives at sea and are excellent underwater swimmers, which is how they catch small fish, their main food
Gentoo Penguin (Pygoscelis papua) albino, Salisbury Island, South Georgia
Galapagos penguins (Spheniscus mendiculus) underwater with shoal of fish, Tagus Cove, Isabela Island, Galapagos
Great frigatebird (Fregata minor) pair, male on left. Galapagos, Ecuador
Gannet (Morus bassanus) Shetland, England, UK. July
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Northern rockhopper penguin (Eudyptes moseleyi) parent with chick. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Emperor penguin (Aptenodytes forsteri) tobogganing, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Royal tern (Sterna maxima) perching on a sand castle on a beach, Fort De Soto Park, St. Petersburg, Florida, USA, March
Crested auklets (Aethia cristatella) pair interacting while perched on rock, St Paul Island, Pribilofs, Alaska, USA, July
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Puffins (Fratercula arctica) and Razorbills (Alca torda) with Eilean Mhuire behind, Shiant Isles, Outer Hebrides, Scotland, UK. June, 2018
King penguins 1+Aptenodytes patagonicus+1 group of three on the shore, St. Andrews Bay, South Georgia. November
King penguins 1+Aptenodytes patagonicus+1. Grytviken, South Georgia. November
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Atlantic puffin (Fratercula arctica) portrait, Machias Seal Island, Maine, USA. July
Waved albatross (Phoebastria irrorata) head portrait, Espanola Island, Galapagos, South America