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Harlequin Shrimp (Hymenocera elegans) on sea floor. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
Two Tawny nurse sharks (Nebrius ferrugineus) on seabed at dusk, Vavuu Atoll, Maldives, Indian Ocean
Kelp greenling (Hexagrammos decagrammus), female, resting on bed of kelp (Laminariales), Prince William Sound, Alaska, USA, Pacific Ocean
Pacific red hermit crab (Elassochirus gilli) on sea floor, Prince William Sound, Alaska, USA, Pacific Ocean
Whitetip reef shark (Triaenodon obesus) swimming in shallows, El Finado, Isabela Island, Galapagos Islands, Ecuador. Pacific Ocean
Painted frogfish (Antennarius pictus) portrait, Sipidan, Malaysia, Celebes Sea
Sea star (Odontaster validus) on Ribbon kelp (Durvillaea antarctica), Antarctic Peninsula, Antarctica, Southern Ocean
Two Speckled moray eels (Gymnothorax dovii) feeding on a Sea turtle carcass on the seabed, Wolf Island, Galapagos Islands, Pacific Ocean
Horned sea star / Chocolate chip sea star (Protoreaster nodosus) on the rocky seabed, Isabela Island, Galapagos Islands, Pacific Ocean
Lemon shark (Negaprion brevirostris) pup and school of fish swimming through Red mangrove (Rhizophora mangle) nursery, Eleuthera Island, Bahamas, North Atlantic
European Spider crab aggregation (Maja squinado) St. Ives, Cornwall. AugustEuropean Spider crab aggregation (Maja squinado) St.Ives, Cornwall. August
Green sea turtle (Chelonia mydas) feeding on Seagrass (Cymodocea nodosa) on the seabed, Tenerife, Canary Islands, Atlantic Ocean
Shoal of fish swimming around an outfall pipe on the seabed discharging treated water, Tenerife, Canary Islands, Atlantic Ocean
Ocellated eagle ray (Aetobatus ocellatus) searching for food in the sand over the seafloor, Mu Koh Similan National Park, Andaman Sea, Phang-nga, Thailand
Gold coral (Savalia Savaglia) colonising Red gorgonian (Paramuricea clavata), Marine Protected area Punta Campanella, Costa Amalfitana, Italy, Tyrrhenian Sea
Streaked gurnard (Trigloporus lastoviza) portrait, Marine Protected area Punta Campanella, Costa Amalfitana, Italy, Tyrrhenian Sea
Splendid toadfish (Sanopus splendidus) emerging from its daytime hiding place at night, Cozumel, Mexico, Caribbean Sea
Great hammerhead shark (Sphyrna mokarran), critically endangered, swimming over sandy seabed accompanied by Blue runner jacks (Caranx crysos) and Remoras (Echeneis sp.), Bimini, Bahamas, Caribbean Sea
Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water with barbels clearly visible on top lip, head portrait, The Bahamas National Shark Sanctuary, South Bimini, Bahamas
Diverse array of algae species including Sargassum weed (Sargassum horneri), Kelp (Laminariales), Sea lettuce (Ulva lactuca) on shallow reef with juvenile Pollock (Pollachius pollachius)
Ancient Roman tessellatum mosaic in black and white decorated with pattern of hexagons, perfectly preserved, in Villa a Protiro. Marine Protected Area of Baia, Naples, Italy. Tyrrhenian sea
Detail of splendid ancient Roman tessellatum mosaic in black and white decorated with pattern of hexagons, perfectly preserved, in Villa a Protiro. Marine Protected Area of Baia, Naples, Italy
Statue depicting companion of Ulysses, perhaps Baio, located in submerged Nymphaeum of Emperor Claudius. Marine Protected Area of Baia, Naples, Italy. Tyrrhenian sea. statue encrusted with sealife
Scuba diver exploring ancient Roman mosaic from the third century AD, with maritime theme, made with black and white tessellatum tiles depicting seabed with dolphins, sea urchins
Scuba diver exploring splendid ancient Roman tessellatum mosaic in black and white decorated with pattern of hexagons, perfectly preserved, in Villa a Protiro
Scuba diver near ancient Roman Statue of Antonia minor, member of Julio-Claudian dynasty, daughter of Marcus Anthony and sister of emperor Augustus, in submerged Nymphaeum of Emperor Claudius
Ancient Roman statue of Antonia minor, member of Julio-Claudian dynasty, daughter of Marcus Anthony and sister of emperor Augustus, located in submerged Nymphaeum of Emperor Claudius
Dungeness crab (Metacarcinus magister) standing on seabed, Vancouver, British Columbia, Canada, Pacific Ocean
Peacock flounder (Bothus lunatus) camouflaged on sand seabed, Dominica, Eastern Caribbean
Male Yellowface pike-blenny (Chaenopsis limbaughi), living in abandoned tubeworm holes in sand, flaring its dorsal fin in threat display, Dominica, Eastern Caribbean
Bobtail squid (Heteroteuthis hawaiiensis) on the seabed, Komodo National Park, Indonesia, Pacific Ocean
Steller sealion (Eumetopias jubatus) swimming over seabed, Vancouver Island, British Columbia, Canada, Pacific Ocean
Pair of mating Big blue octopuses (Octopus cyanea) where the male, on the right, is fertilising the female using its fertilising tentacle. Indian ocean, Maldives
Banded houndshark (Triakis scyllium) group above sea floor. Chiba Prefecture, Honshu, Japan. October
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Redspotted catshark (Schroederichthys chilensis) on sea floor amongst Sea urchins (Echinoidea). Las Tacas, Chile. November
Pacific starry skate (Beringraja stellulata) on sea floor. Vancouver Island, British Columbia, Canada. August
Circular stingaree (Urolophus circularis) above sea floor. Albany, Western Australia. October
Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Yellow seafan (Leptogorgia viminalis) La Gomera, Canary Islands
Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, Bimini, Bahamas
Flag pen shell (Atrina vexillum) in Passe Magnan / Magnan channel, Aldabra, Indian Ocean
Caribbean reef shark (Carcharhinus perezi) cruising over a sand flat in The Bahamas
Light patterns on the seabed and fluffy cumulus clouds in the sky in a split level image. North Sound, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea