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Spotted Porcupine fish (Diodon hystrix) inflated, displaying defensive behaviour, North Pacific Ocean, Hawaii
Oval squid (Sepioteuthis lessoniana) photographed at night, Green Island, Taiwan. Minimum fees apply
Scalloped ribbonfish (Zu cristatus) juvenile, Balayan Bay, Batangas, the Philippines
Galapagos sea lion (Zalophus wollebaeki) inquisitive pup playing with Sally Lightfoot Crab (Grapsus grapsus). Plazas Island, Galapagos, Ecuador
Kelp bass (Paralabrax clathratus) in Giant kelp (Macrocystis pyrifera) forest, Santa Barbara, California, USA
RF - Common hermit crab (Pagurus bernhardus) scuttles over the seabed. Highlands, Scotland, United Kingdom. (This image may be licensed either as rights managed or royalty free.)
RF - Portrait of a coral grouper (Cephalopholis miniata) next to a sea fan (Annella sp.) on a coral reef. Raja Ampat, West Papua, Indonesia. Ceram Sea
RF - Split level photo of mangrove scenery, with hard corals (including Goniopora sp. Porites sp.) growing below Red mangrove tree (Rhizophora sp.). Raja Ampat, West Papua, Indonesia
RF - Blue shark (Prionace glauca) cruises beneath the surface of the English Channel with sunbeams. Cornwall, England, UK. (This image may be licensed either as rights managed or royalty free.)
RF - Grey seal (Halichoerus grypus) relaxing in a bed of kelp (Laminaria digitata), with two seals in the background. Farne Islands, Northumberland, England, United Kingdom. British Isles. North Sea
Queen scallop (Aequipecten opercularis) feeding. Glencoe, Ballachulish, Lochaber, The Highlands, Scotland, United Kingdom. Loch Leven, Loch Linnhe, North East Atlantic Ocean. British Isles
King penguins (Aptenodytes patagonicus) congregate on the beach, seeking a safe point to go into the sea, with male Southern elephant seals (Mirounga leonina) blocking their path
Blue-barred parrotfish (Scarus ghobban), El Pardito Island, Sea of Cortez (Gulf of California), Mexico, September
Swimming crab (Cronius ruber) Canary Islands, Tenerife. Invasive species
Grey triggerfish (Balistes capriscus) portrait, Tenerife, Canary Islands. August
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands. January
Dunstanburgh Castle on headland of the Whin Sill, eroded dolerite boulders in foreground. Northumberland, England, UK. July 2019
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Group of Bigfin reef squids (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Bigfin reef squid (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Brown fur seal / Cape fur seal (Arctocephalus pusillus), Western Cape, South Africa. Atlantic Ocean
Palmers globemallow (Sphaeralcea palmeri) and view towards Toro Islet and Guadalupe Island, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, September
Day octopus (Octopus cyanea) female, in mid-water, Hawaii
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
Queen conch (Aliger gigas) in a seagrass (Thalassia testudinum) meadow, Bahamas
Red sea fan (Paramuricea clavata) with gorgonian corals and Light-bulb sea squirt (Clavelina lepadiformis) Punta Carena, Capri Island, Sorrentine Peninsula, Italy, Tyrrhenian Sea, Mediterranean
Crinoid or feather star (Antedon mediterranea) on Violescent sea whip or Red sea fan (Paramuricea clavata) Punta Campanella Marine Protected Area, Amalfi Coast, Italy, Tyrrhenian Sea, Mediterranean
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
Sea anemone (Calliactis parasitica) usually associated with hermit crabs, here on a Mediterranean bonnet shell (Semicassis granulata), Ponza island, Italy, Tyrrhenian Sea, Mediterranean
Loggerhead turtle (Caretta caretta) accompanied by pilotfish (Naucrates ductor) swimming near the surface. Santa Maria Island, Azores, Portugal, Atlantic Ocean. Vulnerable
Low tide on Westward Ho! beach, tinged pink at sunrise. Devon, England, UK. February 2020
Thrift (Armeria maritima) flowering on cliff top at sunset. Hartland Quay, Devon, England, UK. May 2019
The Green Bridge of Wales sea arch and stack along limestone coastline. Castlemartin, Pembrokeshire, Wales, UK, September 2019
Atlantic spotted dolphins, (Stenella frontalis), Formigas Islet dive site, Santa Maria Island, Azores, Portugal, Atlantic Ocean
Spotted gully shark (Triakis megalopterus) in coral reef. Simons Town, Western Cape, South Africa. June
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Pyjama shark (Poroderma africanum) in reef. Simons Town, Western Cape, South Africa. June
Pyjama shark (Poroderma africanum), two in reef, amongst kelp. Simons Town, Western Cape, South Africa. May
Japanese bullhead shark (Heterodontus japonicus) on reef. Chiba Prefecture, Honshu, Japan. May
Eastern shovelnose ray (Aptychotrema rostrata) on sea floor. Nelson Bay, New South Wales, Australia. July
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Banded sea krait (Laticauda colubrina) hunting in seagrass bed. Manado, North Sulawesi, Indonesia
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
Neptune seagrass (Posidonia oceanica) bed, sun rays shining through water. A patch of seagrass bed in the Mediterranean sea is considered to be the oldest living organism on earth. Spain. June
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Orange and white soft corals, Dead mans fingers (Alcyonium digitatum) thriving beneath kelp (Laminaria hyperborea). St Abbs, Eyemouth, Berwickshire, Scotland, United Kingdom. North Sea