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Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
RF - Lemon shark (Negaprion brevirostris) swimming through sun rays in the late afternoon. Little Bahama Bank, Bahamas. West Atlantic Ocean
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Oman anemonefish / clownfish (Amphiprion omanensis) in a a Bulb-tentacle sea anemone (Entacmaea quadricolor), coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
Blacktip reef shark (Carcharhinus melanopterus) juvenile in shallow water, Maldives, Indian Ocean
Patters of sunlight on a shallow sandy seabed. North Sound, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013
Blue triggerfish (Pseudobalistes fuscus) Egypt, Red Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Bottlenose dolphins (Tursiops truncatus) approaching with curiosity, Dolphin Reef, Eilat, Israel. Red Sea
Sargassum fish (Histrio histrio) near the surface on floating Sargassum seaweed, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Sunset at Trebarwith Strand with incoming tide, North Cornwall, UK. March 2014. *NOT AVAILABLE FOR UK 2016 RETAIL CALENDARS
Detail of rhinopores and the colour pattern of a toxic Nudibranch (Goniobranchus coi) on a coral reef. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Silky shark (Carcharhinus falciformis), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Southern stingray (Dasyatis americana) female swimming over a shallow sand bank with ripples. Stingray City Sandbar, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in their home, a Sea anemone (Heteractis magnifica), on a coral reef. Big Brother island, Red Sea. Egypt
American crocodile (Crocodylus acutus), underwater, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable species
Green turtle (Chelonia mydas) female on a coral reef. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Green sea turtle (Chelonia mydas) female resting on a coral reef. Rock Islands, Palau, Mirconesia. Tropical west Pacific Ocean
Green turtle (Chelonia mydas) with rays of sunlight, Akumal, Caribbean Sea, Mexico, January. Endangered species
Juvenile Green turtle (Chelonia mydas) swimming near the surface, split level view, Fakarava atoll lagoon, Tuamotu Archipelago, French Polynesia, Pacific Ocean
Squid (Abraliopsis atlantica), with photophores in 6 straight longitudinal rows on mantle visible. They are used to produce bioluminescencent light
Juvenile Squid (Onychoteuthis sp. ) deep sea species from Atlantic Ocean off Cape VerdeJuvenile Squid (Onychoteuthis sp.) deep sea species from Atlantic Ocean off Cape Verde. | Captive
Squid (Histioteuthis sp. ) deep sea species from Atlantic Ocean off Cape Verde. CaptiveSquid (Histioteuthis sp.) deep sea species from Atlantic Ocean off Cape Verde. Captive
Johnsons abyssal seadevil, (Melanocetus johnsonii) deep sea species from Atlantic Ocean off Cape Verde. Captive
Common fangtooth (Anoplogaster cornuta) portrait, deep sea species from Atlantic Ocean off Cape Verde. Captive
Ribbon sawtail fish (Idiacanthus fasciola) from Atlantic Ocean, at a depth of 800-1000m
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Giant deep-sea isopod (Bathynomus giganteus) Picture was taken in cooperation with the Zoological Museum University of Hamburg
RF - Portrait of Harp seal (Phoca groenlandicus) pup on sea ice. Magdalen Islands, Gulf of St Lawrence, Quebec, Canada
Sperm whale (Physeter macrocephalus) portrait, with remora fish, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable