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Weaver ants (Oecophylla smaragdina) holding leaves together during nest building, Malaysian Borneo
Weaver ants (Oecophylla smaragdina) portrait, Sabah, Malaysian Borneo
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
Proboscis monkey (Nasalis larvatus), Sabah Malaysia, Borneo
Leaf insect (Phyllium sp. ) camouflaged in rainforest, Mulu National Park, BorneoLeaf insect (Phyllium sp.) camouflaged in rainforest, Mulu National Park, Borneo
Spider hunting wasp (Pompilidae) Maliau Basin, Sabah, Borneo
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
RF - Green sea turtle (Chelonia mydas) beneath clouds. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Cup fungus (Cookeina sp) Danum Valley, Sabah, Borneo
Pink stink bug (Tessaratomidae). Danum Valley, Sabah, Borneo
Red leaf monkey (Presbytis rubicunda) mother and baby, Danum Valley, Sabah, Borneo, Malaysia August
Dawn over the Segama River, with mist hanging over lowland rainforest. Heart of Danum Valley, Sabah, Borneo
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Orange backed red langur (Presbytis rubicunda chrysea) in rainforest understorey. Danum Valley, Sabah, Borneo, Malaysia
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Montane horned frog (Megophrys kobayashii) head close up, Kinabalu National Park, Sabah, Borneo, Malaysia
Minute / Dwarf filefish (Rudarius minutus) just over an inch long as an adult, Celebes Sea, Mabul Island, Malaysia
Sunda leopard cat (Prionailurus javanensis) kitten. Sabah, Malaysian Borneo
Blyths paradise flycatcher (Terpsiphone affinis) male calling, Sabah, Malaysian Borneo
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Lowland dipterocarp rainforest, with huge Tualang (Tapang) or Mengaris Tree (Koompassia excelsa) in the foreground, Danum Valley, Sabah, Borneo
Western / Horsfields tarsier (Tarsius bancanus) hunting invertebrate prey in rainforest understory vegetation at night. Danum Valley, Sabah, Borneo
Giant shield bug nymphs (Pycanum sp. ), recently hatched from eggsGiant shield bug nymphs (Pycanum sp.), recently hatched from eggs. Danum Valley, Sabah, Borneo
Safety in Numbers - Giant Shield bug nymphs (Pycanum sp.), recently hatched from eggs, with a final individual still to emerge
Arboreal tarantula (Lampropelma sp. ) female guarding spiderlings at entrance to hollowArboreal tarantula (Lampropelma sp.) female guarding spiderlings at entrance to hollow retreat, 4 metres up a tree. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of wing. Danum Valley, Sabah, Borneo
Lantern bug (Pyrops whiteheadi), detail of head. Danum Valley, Sabah, Borneo
Lowland dipterocarp rainforest at dawn, Danum Valley, Sabah, Borneo
Sumatran pitviper (Trimeresurus sumatranus), detail of scales. Danum Valley, Sabah, Borneo
RF - Maliau Falls, Maliau River. Centre of Maliau Basin - Sabah, Borneo. (This image may be licensed either as rights managed or royalty free.)
Portrait of Bornean Horned Frog (Megophrys nasuta) amongst leaf-litter in forest floor. Taken at night. Danum Valley, Sabah, Borneo
Flower of locally endemic Rafflesia (Rafflesia tengku-adlinii) flower diameter 22cm blooming on forest floor within lowland Dipterocarp rainforest, Maliau Basin, Sabahs Lost World, Borneo
Maroon / Red leaf monkey / Langur (Presbytis rubicunda) eating fruits in tree, Danum Valley Conservation Area, Sabah, Borneo, Malaysia
Crocodilefish (Cymbacephalus beauforti) close up detail of the retractable eye-lashes, Mabul Island, Sabah, Borneo, Malaysia Celebes Sea
RF - Camouflaged moth (Agathia codina) Kinabalu National Park, Sabah, Malaysian Borneo. September
Moth (Eumelea sp) Danum Valley, Sabah, Borneo
Rhinoceros beetle (Chalcosoma moellenkampi) climbing up tree trunk, Danum Valley Reserve, Sabah, Borneo, Malaysia
RF- Sunrise and mist over lowland dipterocarp rainforest. Danum valley, Sabah, Borneo, Malaysia
Spiky flower-mimic stick insect (Toxodera berieri) active at night. Danum Valley, Sabah, Borneo
Adult Violin Beetle (Mormolyce borneensis) on decaying leaf on the rain forest floor. Near Ginseng Camp in the heart of Maliau Basin - Sabahs Lost World. Maliau Basin, Borneo
Gulik Falls, edge of southern plateau, Maliau Basin, Sabahs Lost World, Borneo (digitally stitched image)
Tropical Butterfly (Adepha) on a leaf with wings spread. Danum Valley, Sabah, Borneo, Malaysia
Eye and spiracle detail of a Bluespotted / Ribbontail Stingray (Taeniura lymma) Sipadan Island, Sabah, Borneo, Malaysia. May
Bearded pig (Sus barbatus) foraging for food at night, Maliau Basin, Sabah, Borneo