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Swallowtail butterflies (Papilio machaon) sunning on a riverbank in Arkhyz section of the Teberdinsky biosphere reserve, Caucasus, Russia, July 2008
Fallen Nordmann fir (Abies nordmanniana) tree, old-growth forest, Arkhyz valley, western part of the Teberdinsky Biosphere reserve, Caucasus, Russia, July 2008
Swallowtail butterfly (Papilio machaon) sunning on a riverbank in Arkhyz section of the Teberdinsky biosphere reserve, Caucasus, Russia, July 2008
First light on snow covered mountain near Dombay, Teberdinsky biosphere reserve, Caucasus, Russia, July 2008
Mountain and glaciers before sunrise, near Dombay, Teberdinsky biosphere reserve, Caucasus, Russia, July 2008
Old male Great rosefinch (Carpodacus rubicilla) on rock, Mount Cheget, Caucasus, Russia, June 2008
Mount Donguzorun (4, 448m) Caucasus, Russia, June 2008
(Anemone sp) flowers, Mount Cheget, Caucasus, Russia, June 2008
Blue flowers (Primula sp) Mount Cheget, Caucasus, Russia, June 2008
Young Baikal seal (Pusa sibirica) at breathing hole. Lake Baikal, Russia, April
Arctic foxes (Vulpes lagopus) juveniles playing, in summer coat, Wrangel Island, Far Eastern Russia, August
Musk ox (Ovibos moschatus) portrait, Wrangel Island, Far Eastern Russia, March
Ice pile of broken shelf ice, near the shore of Lake Baikal, Siberia, Russia, March
Ice formation at sunset, Lake Baikal, Siberia, Russia, March
Red Fox (Vulpes vulpes) female with nest and chicks in mouth. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) portrait, resting on snow. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, walking down a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Mountain Hare (Lepus timidus) feeding. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, Arctic circle Dive Center, White Sea, Karelia, northern Russia. Captive
Grey wolf running silhouetted {Canis lupus} Toropets, Russia
Ice splash out formation with icicles, Lake Baikal, Siberia, Russia, March
Clear panes of ice on Lake Baikal, Siberia, Russia, March
Cracks in the ice of Lake Baikal, Siberia, Russia, March
Large male Reindeer (Rangifer tarandus) standing in shallow waters of the Kronotsky lagoon, away from biting insects. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) jumping across stream. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Kamchatka Brown Bear (Ursus arctos beringianus) stands on its back legs to more easily determine the source of an unfamiliar sound
Ranger station in winter with Kamchatka Brown Bear (Ursus arctos beringianus), Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June. Sequence 3 of 3
Abstract ice formations seen from underwater, Arctic circle Dive Center, White Sea, Karelia, northern Russia, March 2010
Beluga whale (Delphinapterus leucas) swimming under ice, Arctic circle Dive Center, White Sea, Karelia, northern Russia captive
Kamchatka Brown Bear (Ursus arctos beringianus) on the Pacific coast of Kamchatka. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Brown bear {Ursus arctos} near photographers wooden cabin, Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} at the Stained glass window in the Valley of the GeysersBrown bear {Ursus arctos} at the " Stained glass window" in the Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} at the Malachite Grotto, Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} fishing for salmon in the Kronotskaya River at the foot of the Kronotsky Volcano, covered with a fresh coat of snow in September, Kronotsky Zapovednik, Kamchatka
Stellers sea eagles {Haliaeetus pelagicus} gather beside Kuril Lake with Ilinsky volcano in the background, Kamchatka, Far East Russia, winter 2007
Beluga whale (Delphinapterus leucas) play biting camera, Arctic circle Dive Center, White Sea, Karelia, northern Russia. Captive
Brown bear {Ursus arctos} enjoys hot steam from a geyser in Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Ancient burial grounds, Daurian Nature Reserve. Daurian Steppes UNESCO World Heritage Site, Zabaykalsky Krai, Siberia, Russia, June 2016
Traditional mongolian Ovoo shrine / Buddhist shrine near border between Russian and Mongolia. Daurian Nature Reserve. Daurian Steppes UNESCO World Heritage Site, Zabaykalsky Krai, Siberia, Russia
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, with scuba diver in background, Arctic circle Dive Center, White Sea, Karelia, northern Russia, captive
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Brown bear (Ursus arctos) running through water, fishing, Kamchatka, Far east Russia, August
Brown bear (Ursus arctos) walking beside lake, Kamchatka, Far east Russia, August
Alpine meadows with Caucasian scabious (Scabiosa caucasica) in Bezengi gorge, July, 3000m. W-C Caucasus, Kabardino-Balkarskiy nature reserve, Russia
Brown bear (Ursus arctos) standing beside lake looking for salmon, Kamchatka, Far east Russia, August
