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Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April
Black Grouse (Tetrao tetrix) males in winter, Tver, Russia. April
Arctic fox (Vulpes lagopus) in summer moult stealing Snow goose (Chen caerulescens caerulescens) egg, with Snow goose flapping wings cand calling, Wrangel Island, Far Eastern Russia, June
Arctic Fox (Vulpes lagopus semenovi), in dark summer pelage, patrolling cliffside rocks along the coast of Medny Island in the Commander Islands (aka Komandorsky Islands), Russia, July
Male Saiga antelope (Saiga tatarica) running, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009. WWE OUTDOOR EXHIBITION. WWE BOOK
Male European green toad (Bufo viridis) in pond (at 2, 711m) in Adylsu valley, side valley to Baksan valley and Elbrus, Caucasus, Russia, June 2008
(Veronica sp) in flower, Mount Cheget, Caucasus, Russia, June 2008
Transparent ice formation on Lake Baikal, Siberia, Russia, March 2012
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) curled up, with an eye open. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May
Ice fragment reflected in the ice of Lake Baikal, Siberia, Russia, March
Ice fragment on crack in Lake Baikal ice, Siberia, Russia, March
Clear pane of ice, Lake Baikal, Siberia, Russia, March
Portrait of male Reindeer (Rangifer tarandus) with large antlers, in velvet. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June
Ice pile of broken shelf ice, near the shore of Lake Baikal, Siberia, Russia, March
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Old male Great rosefinch (Carpodacus rubicilla) on rock, Mount Cheget, Caucasus, Russia, June 2008
West Caucasian tur {Capra caucasica} with young, Kabardino-Balkarsky NR, Central Caucasus, Russia
Dombai-Ulgen peak with Rhododendron luteum plant, Teberdinskii reserve, NW Caucasus, Russia
Red knot (Calidris canutus rogersi) incubating nest on a coastal gravel spit. Chukotka, Russia. June
RF - Siberian tiger (Panthera tigris altaica) in snow, captive
Footprints of an Amur / Siberian tiger (Panthera tigris altaica) in sand along the edge of the Sea of Japan, Lazovskiy Zapovednik Nature Reserve, Primorsky Krai, Russian Far East. February 1992
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Silhouette of Brown bear (Ursus arctos) beside Lake Kuril at dawn, Kamchatka, Far east Russia, August 2008
Red fox (Vulpes vulpes) cub biting tail of adult male, Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East. June
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, walking down a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Red Fox (Vulpes vulpes) in snowy landscape with Kronotsky Volcano, Zapovednik, Kamchatka, Russia
Red Fox (Vulpes vulpes) stretching. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East
Light reflecting on the surface of Lake Baikal, Siberia, Russia. October, 2021
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) doe with fawn standing on rocky outcrop overlooking forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, on a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012
Snow goose (Chen caerulescens) mobbing Arctic fox (Alopex lagopus) Wrangel Island, Far East Russia, June
Bowhead whale (Balaena mysticetus) tail slapping, Sea of Okhotsk, Russia
Aerial view of Wrangel Bay, Russia, one of the locations where bowhead whales (Balaena mysticetus) of the endangered Sea of Okhotsk subpopulation visit during the summer
Clouds over Lake Baikal, Siberia, Russia, October 2010