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Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Red deer, (Cervus elaphus), stag in falling snow on moorland, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stag in falling snow on moorland, Scotland, UK.February
Reindeer (Rangifer tarandus) female near Aviemore, Cairngorms National Park, Scotland, UK, December
Male and female White-tailed deer (Odocoileus virginianus) with snowman, New York, USA, February
Moose (Alces alces) bull, crossing mountain river after sunset. Grand Teton National Park, Wyoming, USA. October 2017
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Chamois (Rupicapra rupicapra) Ballons des Vosges Regional Natural Park, Vosges Mountains, France, October
Alpine ibex (Capra ibex) on a mountain side in autumn with European larch tree (Larix decidua) trees in the back the Alps, Gran Paradiso National Park, Italy. November
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Red Deer (Cervus elaphus) portrait of head of stag and antlers. Alladale Estate, Scotland, UK, October
Chinese Water Deer (Hydropotes inermis) Norfolk UK September
Red deer (Cervus elaphus) stag roaring during rut, with resting females, England, UK. September
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
Roe deer (Capreolus capreolus) fawn in meadow, Scotland, UK. August
Reindeer (Rangifer tarandus) adult female, antlers in velvet, walking across upland moor, Cairngorms, Scotland, UK, August
Alpine Ibex (Capra ibex) in its landscape, Valsavarenche, Gran Paradiso national park, Aosta Valley, Italy, May
Female Uganda kob (Kobus kob thomasi) suckling calf. Murchisson Falls National Park, Uganda. February
White-lipped deer (Cervus albirostris) stags in mountain landscape, Serxu County, Garze Prefecture, Sichuan Province, China
White-lipped deer (Cervus albirostris) male, Serxu, Garze Prefecture, Sichuan Province, China
Injured Muntjac (Muntiacus reevesi) swimming in a small creek after unsuccessfully attempting to jump across it. This muntjac was unable to get out of the creek himself
Pere Davids deer / Milu (Elaphurus davidianus) during the rutting season, stag, Hubei Tian ezhou Milu National Nature Reserve, Shishou, Hubei, China
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Red deer (Cervus elaphus) stag, Saxony, Germany. September
Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
RF - American Bison (Bison bison) male in snow covered in frost, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Southern lechwe (Kobus leche), two males about to fight, facing each other. Okavango Delta, Botswana
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Himalayan ibex (Capra sibirica) male on snow, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Moose (Alces americanus) close-up grazing, Denali National Park, Alaska, USA, September
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
Mud-covered face of bull African buffalo (Syncerus caffer) with a cateract in one eye, on the banks of the Chobe River, Chobe National Park, Botswana
Roe deer (Capreolus capreolus) young male walking through snow. Close-up, portrait. Norway, February
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Scimitar-horned oryx (Oryx dammah) captive in enclosure of Souss Massa National Park, Morocco. Extinct in the Wild
Chamois (Rupicapra rupicapra) in the snow, Jura Switzerland, November
Alpine ibex (Capra ibex) adult male in deep snow on a ridge with young during snowfall, Gran Paradiso National Park, the Alps, Italy. January
African buffalo (Syncerus caffer) feeding on water lillies in Chobe River, Chobe National Park, Botswana. May
Indian hog deer (Hyelaphus porcinus) mother and fawn, Manas National Park UNESCO World Heritage Site, Assam, India
Blackbuck (Antelope cervicapra), male, Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Red Deer (Cervus elaphus) herd in forest in snow, Cairngorms National Park, Scotland, UK. December
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Alpine ibex (Capra ibex) group posing on a rocky mountain side on misty day, Gran Paradiso National Park, Italy. November
Svalbard reindeer (Rangifer tarandus platyrhynchus) with bloody antlers shedding velvet, Spitsbergen, Svalbard, Norway, September
Marsh deer (Blastocerus dichotomus), stag. Pousada Araras, northern Pantanal, Mato Grosso, Brazil
Red deer (Cervus elaphus) stag bellowing, Los Alcornocales Natural Park, southern Spain, September
